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BugGuide Gathering
Smoky Mountains
University of Tennessee Biological Field Station
August 8-10, 2008
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metamorphosis

Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
No Taxon (Glossary --use Taxonomy, then Info tabs to view entries)
No Taxon (M)
No Taxon metamorphosis
Identification

I-Insects undergoing simple metamorphosis typically have a three-stage life cycle:
1-embryo (egg)
2-nymph (resemble small adults)
3-adult (sexually mature, wings typically functional)

Simple metamorphosis may be further divided into three types:
1-ametabolous development (no metamorphosis). The adults and nymphs are wingless, and there is no visible change in form between the stages, other than in size. Ametabolous development occurs in the Apterogytes and in other groups that undergo simple metamorphosis where the adults are wingless.

2-hemimetabolous development (incomplete metamorphosis). Immature insects (nymphs, often called naiads) are aquatic and breathe via gills. This pattern of development occurs in:
Ephemeroptera (mayflies)
Odonata (dragonflies and damselflies)
Plecoptera (stoneflies)

3-paurometabolous development (gradual metamorphosis). Nymphs resemble small adults and typically have external wing buds. They live in the same habitat as adults, typically taking the same food. Characteristic of many orders, for example:
Hemiptera (true bugs and allies)
Orthoptera (grasshoppers and allies)

II-Insects undergoing complete metamorphosis have a four-stage life cycle:
1-egg (embryo)
2-larva, typically worm-like with several stages (instars)
3-pupa (resting, transformative stage)
4-imago (adult, sexual stage, typically winged)

An insect that undergoes complete metamorphosis is called holometabolous. The following orders have this type of development:
Neuroptera - Antlions, Lacewings and Allies
Coleoptera - Beetles
Hymenoptera - Ants, Bees, Wasps and Sawflies
Trichoptera - Caddisflies
Lepidoptera - Butterflies and Moths
Mecoptera - Scorpionflies, Hangingflies and Allies
Diptera - Flies
Siphonaptera - Fleas
Strepsiptera - Twisted-winged Insects

IIA-certain insects undergo hypermetamorphosis, with an active first-instar larva (planidium). Examples occur among beetles:
Ripiphoridae - Wedge-shaped Beetles
Meloidae - Blister Beetles

III-Several intermediate types of metamorphosis are known, that do not fit neatly into the classification of simple/complete above. These include:
Order Thysanoptera - Thrips
Family Aleyrodidae - Whiteflies (Order Hemiptera)
Males of superfamily Coccoidea - Scales (Order Hemiptera)
Print References
Borror et al., An Introduction to the Study of Insects (2)
Works Cited
1.A Dictionary of Entomology
By George Gordh, David H. Headrick
2.Borror and DeLong's Introduction to the Study of Insects
By Norman F. Johnson, Charles A. Triplehorn