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Subfamily Rutelinae - Shining Leaf Chafers

P. japonica aspect - Popillia japonica Anomala - Anomala lucicola Metalic Yellow ? Beetle - Cotalpa lanigera Hairy beetle - Paracotalpa ursina Andrew's Dune Scarab - Pseudocotalpa andrewsi Beetle - Anomala Small beetle - Strigoderma pygmaea unknown beetle - Anomala ellipses
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Longhorn, Leaf and Snout Beetles)
Superfamily Scarabaeoidea (Scarab, Stag and Bess Beetles)
Family Scarabaeidae (Scarab Beetles)
Subfamily Rutelinae (Shining Leaf Chafers)
Explanation of Names
Author is MacLeay 1819, who originally described as its own family, Rutelidae, now regarded as a subfamily of the Scarabaeidae. Name from genus Rutela (Latreille, 1817), an error for Latin rutila (masculine rutilus), meaning red (1).
Numbers
Worldwide, over 4,100 species.
Identification
Elongated to oval, often shiny, scarabs. Distinctive for having unequal, independently moveable tarsal claws, esp. on hind legs (2). Other typical (but not necessarily diagnostic) characters include:
middle tibiae with two apical spurs
pygidium exposed, not completely covered by elytra
scutellum is exposed, not covered by elytra
antennae with 9-10 segments, last three forming a club

Photos illustrating these characteristics:
Food
Adults feed on foliage and fruit. Larvae feed on roots and decaying vegetation (2).
See Also
Cetoniinae - Fruit and Flower Chafers
Melolonthinae - May Beetles and June Bugs
Print References
The Century Dictionary--entry for Rutela (1)
White, pp. 144-147 (2)
Harpootlian, p. 97 (3)
Dillon, p. 506 (4)
Ratcliffe & Paulsen (5)