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BugGuide Gathering
Smoky Mountains
University of Tennessee Biological Field Station
August 8-10, 2008
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Photos from the last gathering (Minnesota 2007)

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Family Passalidae - Bess Beetles

Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Longhorn, Leaf and Snout Beetles)
Superfamily Scarabaeoidea (Scarab, Stag and Bess Beetles)
Family Passalidae (Bess Beetles)
Other Common Names
Peg Beetles, Betsy Beetles, Patent Leather Beetles
Numbers
However, only two species, both in genus Odontotaenius, are currently known to occur in the United States--the other two species are doubtfully (or formerly?) known from Arizona (Schuster, 1994).
Worldwide, about 500 species are known (Schuster, 2001).
Size
30-40 mm
Identification
Large dark beetles with characteristic body form and lifestyle:
robust body, distinct gap between pronotum and elytra
pronotum grooved in middle
elytra deeply grooved
mandibles robust, toothed, project beyond tip of labrum
antennae have 10 segements, including a club of 3 (typically) to 6 segments
antennal plates (lamellae) cannot be pressed together as in Scarabaeidae and related families.
mentum (lower mouthpart) deeply notched, the notch occupied by the large, horn-like ligula, see Dillon, fig. 438 (2)
Each tarsus has five segments (tarsal formula 5-5-5)
Some images of the widespread Horned Passalus, Odontotaenius disjunctus, illustrating these characters:

Tarsi:
Range
Eastern and south-central North America. The Horned Passalus, Odontotaenius disjunctus is widespread in east. Odontotaenius floridanus is a Florida endemic, described in 1994. These are apparently the only two species of the family known to occur in North America (United States and Canada).
Family is cosmopolitan, with most species in the tropics.
Habitat
Live inside rotting logs in forests.
Food
Adults and larvae feed on wood
Life Cycle
Unique subsocial lifestyle. Adults and larvae live together in family groups in galleries excavated in rotting wood by adults. Adults care for larvae, and actively feed them prechewed food. Both adults and larvae stridulate, which is used for communication within the group. See Generic Guide to New World Scarab Beetles for more details.
Print References
Dillon and Dillon (2)
America Beetles, Vol. 2, Chapter 25 (3)
Gordh, p. 670 (4)
Schuster, 1994. Odontotaenius floridanus New Species (Coleoptera: Passalidae): A Second U.S. Passalid Beetle. Florida Entomologist 77(4): 474-478, available here.
Works Cited
1.The Century Dictionary: an encyclopedic lexicon of the English language
2.A Manual of Common Beetles of Eastern North America
By Dillon, Elizabeth S., and Dillon, Lawrence
3.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
By Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.)
4.A Dictionary of Entomology
By George Gordh, David H. Headrick