This species takes two years to mature. The adults of the oak twig pruner fly about the time the oak leaves are beginning to form, and deposit eggs near the tips of twigs. The young larvae mine down the stems, increasing in size, until late summer. They spend the first winter within the twig usually near a node or thickened area. They resume feeding the second season and soon begin their pruning cuts beneath the bark. Larvae cut around the branch except for the thin bark so that the branches break with the wind and fall to the ground. The larvae remain in the fallen branches for up to several months, pupating in the fall. Adults emerge the following spring. Damage is most noticeable in alternate years.
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