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Species Cybister fimbriolatus

Fringed Diving Beetle - Cybister fimbriolatus - male large Dytiscus, ventral view - Cybister fimbriolatus A beetle out of water - Cybister fimbriolatus Cybister fimbriolatus Predaceous Diving Beetle - Cybister fimbriolatus Predaceous Diving Beetle - Cybister fimbriolatus - male diving beetle - Cybister fimbriolatus Cybister fimbriolatus - female
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Adephaga (Ground and Water Beetles)
Family Dytiscidae (Predaceous Diving Beetles)
Subfamily Dytiscinae
Tribe Cybistrini
Genus Cybister
Species fimbriolatus (Cybister fimbriolatus)
Other Common Names
Giant Diving Beetle (1)
"Fringed Diving Beetle" suggested here
Synonyms and other taxonomic changes
Current: Cybister fimbriolatus (Say)
Orig. Comb: Dytiscus fimbriolatus Say 1823
Syn: Cybister fimbriolata--a misspelling sometimes seen or suggested(!) in Internet searches
Explanation of Names
Species name from Latin, fimbria fringed. Fimbriate in entomology refers to a "structure displaying marginal setae of irregular length". (2) This refers to the metatarsi, presumably.
Numbers
Two subspecies are noted in some sources:
Cybister fimbriolatus fimbriolatus (Say, 1825)
Cybister fimbriolatus crotchi Wilke 1920
Ciegler (3) notes that it is difficult to separate these two forms, and that they occur together in South Carolina.
Size
26-33 mm (large)
Identification
Large diving beetle, elytra with yellow margin. Similar to Dytiscus species, but metatarsal claws different. (See guide for genus.) Elytra and pronotum smooth in male. Unique structure on male protarsi, used for grasping female during mating. Dytiscus male has similar structure, but differing in details--two large disks are obvious. White, fig. 33, and Papp (1), figs. 178, 179 illustrate this well (4). Female Cybister has fine furrows on pronotum. Male Cybister has, apparently, two rows of punctures on each elytron, lacking in Dytiscus (4).

Blatchley (5) gives the following description of this species:
Ovate, more or less wedge-shaped. Above brown with a faint greenish tinge; thorax and elytra with a broad yellow margin; front of head, four front legs and spots at sides of abdominal segments 3 to 6, also yellow. Thorax and elytra of female, except along the suture, with numerous fine short impressed lines. Length 30-33 mm. Throughout the State (Indiana); frequent. May 27-September 20. In autumn sometimes found beneath rubbish on the beaches of lakes.
Range
TX-FL-NY-ND / adj Can. (3), (6)
Only member of its genus in southeast.
Habitat
Aquatic: ponds, usually deep water. Comes to lights.
Season
Much of year in south.
February-September, November-December (South Carolina).
March, May, July, September-October (North Carolina).
Food
both larva and adult prey on aquatic insects, fish, etc.
Life Cycle
Seems to have two periods of activity per year, based on reported collection dates.
Remarks
This species attracts attention due to its large size and habit of coming to lights.
See Also
Dytiscus species
compare also Hydrophilus, a Giant Water Scavenger Beetle
Print References
Blatchley, p. 236 (5)
Brimley, p. 132 (7)
Ciegler, p. 88, figs. 5, 102 describes and illustrates C. fimbriolatus. (3)
Dillon and Dillon, pp. 153-154, Plate XVI (8)
Evans, plate 31--color photo of specimen (9)
Gordh, entry for fimbriate (2)
Papp, p. 62, figs. 177-178 (1)
Salsbury, p. 170--photo (10)
Swan and Papp, p. 351, fig. 595--also shows male tarsus (11)
White, fig. 33--illustration of male protarsus, comparing with Dytiscus (4)
Internet References
North Carolina State University lists only C. fimbriolatus from that state--59 pinned.
Clemson University--Cybister sp.
Works Cited
1.Introduction to North American Beetles
By Charles S. Papp
2.A Dictionary of Entomology
By George Gordh, David H. Headrick
3.Water Beetles of South Carolina
By Janet Ciegler
4.Peterson Field Guides: Beetles
By Richard E. White
5.An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana.
By Blatchley, W. S.
6.Predaceous Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska
By D.J. Larson, Y. Alarie, and R.E. Roughley
7.National Audubon Society Field Guide to Insects and Spiders
By Lorus and Margery Milne
8.A Manual of Common Beetles of Eastern North America
By Dillon, Elizabeth S., and Dillon, Lawrence
9.An Inordinate Fondness for Beetles
By Arthur V. Evans, Charles L. Bellamy, Lisa Charles Watson
10.Insects in Kansas
By Glenn A. Salsbury and Stephan C. White
11.The Common Insects of North America
By Lester A. Swan, Charles S. Papp