This group contains the most "primitive" fly families, and is paraphyletic to the rest of the flies. It contains the very diverse crane fly families (Tipuloidea
1,
2,
3), along with several families of blood-feeding flies, such as the mosquitoes (
Culicidae), black flies (Simuliidae), and no-see-ums (Ceratopogonidae). Infraorder arrangement based on Tree of Life.