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Family Tenebrionidae - Darkling Beetles

Beetle - Hymenorus Eleodes parvicollis a load of Corticeus - Corticeus parallelus Long-jointed - Statira gagatina J larvae adult - Mycetochara binotata Eleodes - Eleodes longicollis Eusattus sp.? - Eusattus Beetle on kitchen carpet - Neatus tenebroides
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Longhorn, Leaf and Snout Beetles)
Superfamily Tenebrionoidea (Fungus, Bark, Darkling and Blister Beetles)
Family Tenebrionidae (Darkling Beetles)
Synonyms and other taxonomic changes
Tenebrionidae Latreille, 1802. Includes the Alleculidae (Comb-clawed Beetles) and Lagriidae (Long-jointed Beetles), often considered at Family rank.
Explanation of Names
Tenebrionidae is from genus Tenebrio, that Latin "one who loves darkness, a trickster". In turn tenebrio is from Latin tenebrae, darkness (1) (2).
Darkling as an adjective means "darkening, almost dark, obscure", or "blinded" (Internet searches). An example: "Even in this darkling hour I have faith that this will go on" (Winston Churchill, at a 1949 MIT graduation).
Numbers
About 1200 spp. in the U.S. and Canada grouped in 191 genera.(3)
Size
2-35 mm
Identification
Usually dark, a few colored and/or patterned, sometimes with red. Body shape variable--elongated to more oval, usually flattened. Many large species are flightless and have fused elytra. Characteristics of Tenebrionidae:
first abdominal sternite entire, not divided by hind coxae (characteristic of suborder Polyphaga)--compare Carabidae in the suborder Adephaga, which have that sternite divided
eyes usually notched
antennae variable--thread-like (filiform), bead-like (moniliform), or clubbed
antennae typically with 11 segments, insertion concealed from above
tarsi 5-5-4 (hind tarsi with 4 segments), compare Carabidae, which have a formula of 5-5-5
Larvae are cylindrical and hard-bodied, called "false wireworms" because they resemble larvae of click beetles, Elateridae.
Range
Widespread, but most diverse in the western United States. White (4) says about 140 species found in east, 1,100 found in west.
Habitat
Typically found under stones, decaying logs, bark. A few species diurnal, found in open. Many species are adapted to desert conditions.
Food
Many are scavengers of plant material as both adults and larvae. Some attracted to carrion, dead insects, dung. Some feed on fungus, often found under bark. Some are pests of stored grain and of insect collections.
Life Cycle
Most larvae live in decaying matter. Many species have chemical defenses.
See Also
Ground Beetles - Carabidae
Pleasing Fungus Beetles - Erotylidae
Print References
Arnett, American Beetles, Vol. 2, Chapter 106 (3)
Borror, entry for tenebrio (1)
Dillon, Manual of Common Beetles..., p. 463 (5)
White, Field Guide to the Beetles (4)
The Century Dictionary--entries for tenebrio, tenebrionidae (2)