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Calendar
BugGuide Gathering
Smoky Mountains
University of Tennessee Biological Field Station
August 8-10, 2008
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Photos from the last gathering (Minnesota 2007)

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Genus Brachinus - Bombardier Beetle

Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Adephaga (Ground and Water Beetles)
Family Carabidae (Ground Beetles)
Subfamily Brachininae
Tribe Brachinini
Genus Brachinus (Bombardier Beetle)
Numbers
Genus has forty to fifty North American species--see print references cited below.
Size
5-13 mm
Habitat
Found under loose bark, rocks, boards, etc. Also found on ground in open at night. Usually associated with floodplains, edges of temporary ponds.
Season
Typically spring, early summer and fall? March-July, September-October (B. alternans, South Carolina).
Life Cycle
Eggs laid singly in mud cells made on plants, rocks. Larvae are external parasites of dytiscid beetles such as Hydrophilus. Larvae also scavenge body of host after its death. A few species parasitize whirligig beetles, Gyrinidae. Larvae have reduced legs, typical of parasitic forms. Pupation occurs inside the body of the host. Adults have impressive chemical defenses, ejecting toxic, foul-smelling gases from their abdomen with a loud popping sound. The explosive brew is composed of hydrogen peroxide, hydroquinone, and catalytic enzymes.
Print References
Sikes, pp. 76-77, lists 12 species for Rhode Island (2)
Dillon, pp. 107-108, plate XI, describes and illustrates two species (3).
Papp, p. 51, fig. 141--B. americanus (4)
Arnett and Jacques, p. 98, fig. 254 (5)
White, p. 94, fig. 31 (6)
Milne, p. 537, fig. 166 (7)
Internet References
National Center for Science Education--discussion of mechanism of chemical defenses
Clemson University--discussion of chemical defenses under account for Galerita