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Family Membracidae - Treehoppers

Treehopper - Micrutalis parva Ceresa species - Ceresa albescens Nymphs on Viburnum - Enchenopa onviburnum Homoptera - Cyrtolobus vau Treehoppers:  Adults & Nymphs - Entylia Treehopper - Ophiderma - female Hopper - Cyrtolobus auroreus Treehopper with a Mullet - Micrutalis
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Hemiptera (True Bugs, Cicadas, Hoppers, Aphids and Allies)
Suborder Auchenorrhyncha (Free-living Hemipterans)
Superfamily Cicadoidea
Family Membracidae (Treehoppers)
Other Common Names
Thorn Bugs
Pronunciation
mem-BRAY-ci-dee
Explanation of Names
Family name Membracidae is from genus Membracis Fabricius, 1775. That is New Latin, from Greek membrax (μεμβραξ), a type of cicada (1). (Apparently Latin membrana, source of English membrane, is not related.)
Numbers
60 genera and 257 species in North America according to Arnett (2)
worldwide, at least 600 genera, 3,200 species (Wikipedia--Treehopper)
Size
range 2-20 mm, with most species under 12 mm (2)
Identification
Characters of Membracidae, as well as other families of the Cicadoidea, are (Folz, 2001):
antennae short and bristle-like (aristate), arising in front of or between the compound eyes
tarsi with three segments
middle coxae short and contiguous
no Y-vein in the anal area of the forewing
The Membracidae may be separated from related families by noting the large pronotum that extends back over the abdomen and (often) covers the head:
   
Many species appear humpbacked; others have spines, horns or keels. Some look like thorns.

Food
Adults and nymphs suck plant juices. Many species feed on trees and shrubs, but some species are found on herbaceous plants. Most species feed only on specific types of host plants.
Life Cycle
Eggs overwinter and hatch in the spring. Nymphs drop to the ground. They return to trees to lay eggs. Most species are solitary, but many occur in groups or clusters and exhibit presocial behavior--adults often stay near nymphs, tending them. Many have mutualistic relationships with ants--the ants fend off predators and/or parasites and collect sugary fluid secreted by the hoppers.
Remarks
Only a few species are considered pests; most of the damage is caused by egg laying.
Print References
Arnett, American Insects, pp. 304-306 (2)
Borror and DeLong (3)
The Century Dictionary, entry for Membracis (1)
Internet References
Folz, John (2001). ENY 3005 Family Identification--Homoptera: Cicadoidea: Membracidae (University of Florida, accessed 19 October 2009)
C. H. Dietrich, Section for Biodiversity, Illinois Natural History Survey--Treehoppers
Tree of Life Web--Membracidae
Wikipedia--Treehopper