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Family Coccinellidae - Lady Beetles

h.parenthesis - Hippodamia parenthesis Coleomegilla maculata Black ladybeetle - Axion tripustulatum Coccinellid ? - Brumus quadripustulatus ladybug - Cycloneda munda seven-spotted ladybug - Coccinella septempunctata? - Coccinella septempunctata Different Lady Beetle - Hippodamia sinuata Seven-spotted Lady Beetle? - Coccinella septempunctata
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Longhorn, Leaf and Snout Beetles)
Superfamily Cucujoidea (Flower, Flat Bark and Ladybird Beetles)
Family Coccinellidae (Lady Beetles)
Other Common Names
Ladybird (especially in Canada)
Ladybug
Ladybeetle
Pronunciation
käksə′nelə′dē (kox-ih-NELL-a-DEE)
Synonyms and other taxonomic changes
Coccinellidae Latreille, 1807
Explanation of Names
A group with many interesting biological, cultural, and linguistic associations:
Scientific name: from New Latin coccinella, the diminutive of Latin coccinus, "scarlet." That from Greek kokkinos (κοκκινος), from kokkos, "berry." A related Greek term is kermes, an insect that was used to make a scarlet dye. Compare cochineal, another red insect, and also the bacterium Coccus, from the same root (Internet searches), (1).
Common name: "Ladybird" was first used in medieval England, perhaps because these beneficial predators of agricultural pests were believed to be a gift from the Virgin Mary - the "Lady." Other European names have similar associations, such as the German Marienkafer, "Marybeetle." This may represent an even older, widespread pre-Christian mythology associated with the beetle (Univ. Florida, Wikipedia, Internet searches).
A well-known use of the common name is the popular nursery rhyme Ladybird, Ladybird, fly away home....
Numbers
60 genera and 481 species in America north of Mexico, according to American Beetles.(2)


Family Coccinellidae Latreille, 1807
(adapted from American Beetles)

Subfamily STICHOLOTIDINAE Weiss, 1901
Very small (1-1.5mm), solid black. Strongly convex (dome-shaped), appearing hemispherical to spherical. Antennae and palpi (mouthparts) with distinctive shape.
Tribe Microweisini Leng, 1920
Genus Microweisea Cockerell, 1903

Genus Coccidophilus Brethes, 1905
Genus Gnathoweisea Gordon, 1970
Genus Nipus Casey, 1899

Tribe SERANGIINI Pope, 1962
Genus Delphastus Casey, 1899

Tribe CEPHALOSCYMNINI Gordon, 1985
Genus Cephaloscymnus Crotch, 1873

Subfamily SCYMNINAE Mulsant, 1846
Adults: Usually small, oval to oblong, highly convex (dome-shaped). Larvae: usually covered with white wax that may form long "hairy" tufts.
Tribe SCYMNILLINI Casey, 1899
Small, black, sometimes with red spots. Dorsal surface smooth, pubescent (having short, fine hairs), or both. Legs dark.
Genus Zilus Mulsant, 1850

Genus Zagloba Casey, 1899


Tribe STETHORINI Dobzhansky, 1924
Small, dark brown or black, dorsal surface pubescent. Legs yellow or brown.
Genus Stethorus Weiss, 1885


Tribe SCYMNINI Mulsant, 1846
Round, oval, or oblong. Brown or black; may have brown, yellow, or red markings. Legs yellow to brown except Cryptolaemus with black legs. Dorsal surface and eye pubescent. Larvae: covered in waxy white tufts, resembling mealybugs or hairy caterpillars.

Genus Nephaspis Casey, 1899

Genus Cryptolaemus Mulsant, 1853

Genus Didion Casey, 1899

Genus Scymnus Kugelann, 1794

Genus Sasajiscymnus Vandenberg, 2004
Genus Nephus Mulsant, 1846

Genus Scymnobius Casey, 1899

Tribe DIOMINI Gordon, 1899
Genus Diomus Mulsant, 1850

Genus Decadiomus Chapin, 1933

Tribe SELVADIINI Gordon, 1985
Genus Selvadius Casey, 1899

Tribe HYPERASPIDINI Mulsant, 1846
Round, oval, or oblong. Prominent scutellum (triangle at base of wings). Blue eyes. Bold markings, often colorful: black or dark brown with contrasting spots, stripes, wavy lines. One species metallic blue.
Genus Blaisdelliana Gordon, 1970
Genus Helesius Casey, 1899
Genus Thalassa Mulsant, 1850

Genus Hyperaspis Redtenbacher, 1844

Genus Hyperaspidius Crotch, 1873


Tribe BRACHIACANTHINI Mulsant, 1850
Oval or oblong. Blue eyes, head often with white "face." Dark with white, yellow, orange, or red markings (spots, triangles, diamonds); sometimes yellow, orange or red with black markings.
Genus Brachiacantha Dejean, 1837


Tribe CRYPTOGNATHINI Mulsant, 1850
Genus Cryptognatha, Mulsant, 1850

Subfamily CHILOCORINAE Mulsant, 1846
Round, oval, or shield-shaped. Highly convex, explanate (helmet-like, with a flared "rim"). Often black with 2-6 red or orange spots; sometimes red or orange with or without spots; one species blue. Larvae: usually black, sometimes paler, always with long "spines."

Tribe CHILOCORINI Mulsant 1846
Genus Brumoides Chapin, 1965

Genus Brumus Mulsant, 1850
Genus Axion Mulsant, 1850

Genus Curinus Mulsant, 1850

Genus Arawana Leng, 1908
Genus Exochomus Redtenbacher, 1843

Genus Halmus Mulsant, 1850
Genus Chilocorus Leach, 1815


Subfamily COCCIDULINAE Mulsant, 1846
Tribe COCCIDULINI Mulsant, 1846
Genus Coccidula Kugelann, 1798

Genus Rhyzobius Stephens, 1829

Tribe NOVIINI Mulsant, 1850
Genus Rodolia Mulsant, 1850

Genus Anovia Casey, 1920


Tribe EXOPLECTRINI Crotch, 1874
Genus Exoplectra Chevrolat, 1837

Tribe AZYINI Mulsant, 1850
Genus Azya Mulsant, 1850
Genus Pseudoazya Gordon 1980

Subfamily COCCINELLINAE Latreille, 1807
Many of the most familiar lady beetles. Oval or oblong, sometimes tapering to a point. Usually brightly colored with black markings; or black with colorful markings. A few species white or gray with black markings. Larvae: Dark, "alligator-like," often with colorful markings.

Tribe COCCINELLINI Latreille, 1807
Genus Paranaemia Casey, 1899

Genus Naemia Mulsant, 1850

Genus Coleomegilla Timberlake, 1920

Genus Ceratomegilla Crotch, 1873
Genus Hippodamia Dejean, 1837

Genus Anisosticta Dejean, 1837

Genus Macronaemia Casey, 1899

Genus Aphidecta Weise, 1899
Genus Adalia Mulsant, 1850

Genus Coccinella Linnaeus, 1758

Genus Cycloneda Crotch, 1871

Genus Harmonia Mulsant, 1850

Genus Anatis Mulsant, 1850

Genus Myzia Mulsant, 1846

Genus Calvia Mulsant, 1850

Genus Propylea Mulsant, 1846

Genus Coelophora Mulsant, 1850

Genus Olla Casey, 1899

Genus Neoharmonia Crotch, 1871

Genus Mulsantina Weise, 1906


Tribe HALYZIINI Mulsant, 1846
Small to very small (1-3 mm). Pronotum transparent. White or light brown with brown, black, and sometimes orange spots. Larvae: light gray, may have black and orange markings.

Genus Psyllobora Dejean, 1836

Subfamily EPILACHNINAE Ganglbauer, 1899
Eat plants, unlike most other lady beetles. Adults: Round or oval, highly convex, mouthparts adapted for chewing plants. Orange or yellow with dark spots. Larvae: yellow with long spines.

Tribe EPILACHNINI Costa, 1849
Genus Epilachna Dejean, 1837

Genus Subcoccinella Huber, 1842
Size
From under 1 mm to 10 mm. Larvae from under 1 mm to 12 mm.
Identification
Red lady beetles with black spots are among the most familiar and easily-recognized insects. There are many other colors and patterns, though, which can be hard to identify. The following physical features will identify all lady beetles, no matter what color they are.

Rounded or oval, dorsally convex (dome-shaped), nearly flat on the ventral (bottom)

Pronotum (hard shell in front of wings) often conceals the head from above, and may even look like a head. Photos show the same species with and without head concealed.

Antennae short or very short with 8-11 segments; last 3-6 segments form a weak club. Click the thumbnails for a close look at the details.

Tarsal formula 4-4-4, but appears to be 3-3-3. In other words, each lower leg appears to have three segments.

First abdominal sternite entire, not divided by hind coxae (characteristic of suborder Polyphaga of the Coleoptera)


Larvae
Larvae are very different-looking, and may not be recognized as lady beetles. They are more flattened and elongated ("alligator-shaped"), without the wings to make a round, glossy shell. Predatory species have fully-developed legs, so they don't look as much like worms as many other beetle larvae. Many are gray or black and may have colorful markings (A). Others have waxy white tufts, resembling caterpillars or mealybugs (B). Plant-eating larvae are yellow with long spines (C).
A. B. C.
Range
World-wide, including all of North America.
Habitat
Anywhere with suitable food. Usually outdoors, but in cold weather large groups may enter houses.
Season
Adults and larvae are present from spring to fall, with some adults able to survive cold winters. Year-round in the south.
Food
Most lady beetles are predatory, eating a wide variety of other insects. Prey includes aphids (A, B), mealybugs, scale insects, fly larvae (C), and small caterpillars. They also eat insect eggs (D) and pupae (E) - even those of other lady beetles (F).
A. B. C.
D. E. F.

Some lady beetles are plant-eating, however, and can be destructive in gardens (A). Others consume fungus, such as mildew and plant molds (B). Predatory species may supplement their diet with pollen (C).
A. B. C.
Life Cycle
The larvae are different from the adults - Some have tufts of white wax on the surface that make them look like mealybugs.

Their food is usually similar to that of the adults, and in the predatory species they're often important as pest controls

Before becoming adults, they go through an immobile stage with a shell-like skin. They shed this pupal case to emerge as adults.

see BugGuide Eggs, Larvae, and Pupa info page for more information including current images of species.
Remarks
Identification within the family (at least for adults) often depends on patterns in the pronotum (the area in front of the wings/"shell", but behind the head). A view of your ladybird from the front will help with ID.

Some lady beetles, particularly Harmonia axyridis, serve as hosts to parasitic Laboubeniales fungi, resulting in this appearance (click on pictures for more comments):
See Also
Most other beetles have longer antennae, but some are superficially similar:
Leaf beetles of family Chrysomelidae. Tarsal formula appears 4-4-4, while lady beetles appear 3-3-3.
Endomychidae - Handsome Fungus Beetles
Erotylidae - Pleasing Fungus Beetles
Print References
The Century Dictionary--entry for Coccinella (1)
Castner, p. 120 (3)
White (4)
Larval key to Genera and selected Species - Rees, B.E. (5)
American Beetles, Vol. 2, Chapter 93 (2)
Internet References
Lady Beetles of Ontario images and info (Steve Marshall, U. of Guelph)
Univ. Florida ENY 3005 Family Identification--Coccinellidae
Wikipedia--good reference on cultural associations
Ladybird Morphology Interactive diagram of lady beetle anatomy.
Univ Minnesota Ladybird homepage with keys and photos to common species (including larvae and pupae)
Gordon, Robert D. The Coccinellidae (Coleoptera) of America North of Mexico. Journal of the New York Entomological Society 93(1):1-912.