Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada
Clickable Guide
Moths Butterflies Flies Caterpillars Flies Dragonflies Flies Mantids Cockroaches Bees and Wasps Walkingsticks Earwigs Ants Termites Hoppers and Kin Hoppers and Kin Beetles True Bugs Fleas Grasshoppers and Kin Ticks Spiders Scorpions Centipedes Millipedes


TaxonomyBrowse
Info
ImagesLinksBooksData

Family Carabidae - Ground Beetles

White-cloaked Tiger Beetle - Eunota togata Amblycheila baroni Carabids - Pterostichus corvinus Notiobia sp. - Notiobia - female Carabid - Elaphrus olivaceus Tachys  sp. 2 - Tachys Unknown - Acupalpus testaceus Bembidion inaequale? - Bembidion inaequale
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Adephaga (Ground and Water Beetles)
Family Carabidae (Ground Beetles)
Synonyms and other taxonomic changes
Carabidae Latreille 1802

Includes tiger beetles (Cicindelinae), often treated as a separate family in the past; a recent trend (not followed here) is to treat Rhysodidae within the Carabidae, too
Numbers
by far the largest family of Adephaga and one of the largest insect families with over 30,000 spp. worldwide(1); ca. 2700 spp. in almost 200 genera (assigned to 14 subfamilies and 47 tribes) in our area(2); 930 spp. occur in Canada(3)
sample local faunas: WI 489 spp.(4), GA 531, AL 345, FL 420(5), SC 479(6), NH ~430(7), OK 354(8), CA >700(9)...
Overview of our fauna:
Family Carabidae
Taxa not yet in the guide: native (*), non-native (+); most names are linked to images available on other sites.
Subfamily Paussinae
Subfamily Gehringiinae
Subfamily Omophroninae
Subfamily Nebriinae
Subfamily Carabinae
Subfamily Cicindelinae
Subfamily Loricerinae
Subfamily Elaphrinae
Subfamily Promecognathinae
Subfamily Scaritinae
Subfamily Broscinae
Subfamily Trechinae
Subfamily Harpalinae
Tribe Harpalini
   Subtribe *Pelmatellina *Pelmatellus
Tribe Licinini
   Subtribe Dicaelina Diplocheila, Dicaelus
   Subtribe Licinina Badister
Tribe Lebiini
   Subtribe Apenina Apenes
   Subtribe Cymindidina Cymindis
   Subtribe Lebiina Lebia, Hyboptera
   Subtribe Nemotarsina Nemotarsus
   Subtribe Metallicina Euproctinus
   Subtribe Agrina Agra
Subfamily Pseudomorphinae
Subfamily Brachininae
Size
0.7-66 mm
Identification
Keys to taxa can be found in:
American Beetles (2) (keys to NA genera)
Ground Beetles and Wrinkled Bark Beetles of South Carolina (6) (SC, good for southeast fauna as well)
The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska, parts 1—6 (10) (focuses more on northern fauna)
Florida ground beetles (5) (key to genera and species of a few genera in Florida)
Illustrated Identification Guide to Adults and Larvae of Northeastern North American Ground Beetles (11) (northeastern fauna)
The Beetles of Northeastern North America (12) (older with many errors, but still useful for northeastern fauna)
The Beetles of the Pacific Northwest Pt 1 (13) (northwestern fauna)

Some European sites give a good idea of Holarctic carabid diversity at a glance, e.g.:
Die Käfer-Fauna Deutschlands, by Christoph Benisch
Representative galleries of NA species are provided in(3)(5)(1)(11)
Distinguishing carabid from staphylinid larvae
If you have a larva in hand (well, with some magnification, probably), you can tell carabids from staphs because the former have 6-segmented legs and often 2 claws, while staphs have only 5-segmented legs and always only 1 claw. Also, nearly all carabids have the urgomphi solidly attached to segment 9 (no joint at the base), and at least some of the ones that do have them articulated basally have more than 2 segments, which staphs never have. Staphs almost always have the urogomphi articulated and they have only one or two segments; the ones with solid urogomphi are all little guys (including pselaphines) and quite different in form from carabid larvae. (Margaret Thayer, pers. comm. to Jim McClarin; also see Margaret's comment)
Range
worldwide
Food
Most adults rapidly pursue their prey (other insects) at night. A few eat pollen, berries, and seeds. Most larvae are predators, but some are herbivores or parasitoids.
Remarks
adventive elements of Canadian fauna reviewed in(14)
See Also
Darkling Beetles (Tenebrionidae)

Other beetles superficially resembling carabids:
Print References
Catalogue in (15)
Internet References