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Subfamily Alysiinae

Braconid Wasp Wasp Alysiinae? - Oenonogastra Braconid - female Braconid ex agromyzid ex Pseudelephantopus spicatus Alysiine? ex Pegomya cf. pseudobicolor - Orthostigma Braconid Wasp - male Braconid Wasp ?
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hymenoptera (Ants, Bees, Wasps and Sawflies)
No Taxon ("Parasitica" (parasitic Apocrita))
Superfamily Ichneumonoidea (Braconids and Ichneumons)
Family Braconidae (Braconid Wasps)
Subfamily Alysiinae
Other Common Names
jaw wasps (1)
Pronunciation
Ah-liss-ee-EYE-nee
Numbers
Over 1,000 species worldwide.(2)
Size
generally less than 5 mm; often 2-3 mm.
Identification
Very large marginal cell is the hallmark for several genera in this subfamily.
The vertex of head has a unique shape.
An interesting feature of this subfamily is that the mandibular action is opposite to other wasps. To visualize, place the backs of your hands together and then move your fingers outward.

Comment from Joseph Fortier on the second of the above images: "What gives this braconid away as an alysiine is the last (closeup) shot, showing the truncated, outwardly curving mandible. Alysiine wing venation is often closely similar to that of Opiinae, another fly parasitoid. Alysiines use these specialized mandibles for digging out of the tough fly puparium after emergence from their own pupae."
Range
Worldwide.
Food
Adults feed on pollen and nectar. Larvae feed on immature cyclorrhaphous fly life stages.
Life Cycle
Female alysiine wasps lay their eggs in the eggs or maggots of their hosts, and emerge from the host puparium after eating and killing the host. Thus they are "egg-pupal" or "larval-pupal" parasitoids. The hosts are cyclorraphous Diptera, which are flies with short antennae in which the adult emerges through the hard puparium through a circular opening. In the case of a parasitized fly puparium, the wasp emerges instead of the adult fly.(2) For more info on parasitoid lifeways Click here.
Remarks
Parasitoids of cyclorrhaphous Diptera.
See Also
Compare mandibles of Opiinae:
Print References
Wharton, R.A. Subfamily Alysiinae. In: R.A. Wharton, P.M. Marsh, and M.J. Sharkey. 1997. Manual of the New World Genera of the Family Braconidae (Hymenoptera). (2)
Works Cited
1.Three new species of Chorebus from Spain (Hymenoptera: Braconidae: Alysiinae).
Docavo, I., J. Tormos, and M. Fischer. . 2002. Florida Entomologist.
2.Manual of the New World Genera of the Family Braconidae (Hymenoptera)
Wharton, R.A., P.M. Marsh, M.J. Sharkey (Eds). 1997. International Society of Hymenopterists.