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Subfamily Scarabaeinae - Dung Beetles

Canthon imitator Brown - Canthon imitator Horned Beetle - Liatongus californicus - male BeetlesDungForWeb_9666 - Canthon Onthophagus orpheus, probably - Onthophagus orpheus Canthon indigaceus scarab, at lights at night - Copris Scarabaeidae, dorsal - Onthophagus hecate - female Scarab Beetle - Copris fricator
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga
Superfamily Scarabaeoidea
Family Scarabaeidae (Scarab Beetles)
Subfamily Scarabaeinae (Dung Beetles)
Synonyms and other taxonomic changes
see (1)
Numbers
б,850 spp. in ~280 genera worldwide(2)
Identification
keys to MD spp. in (3)
Tribe ID detail
Works Cited
1.The taxonomic revolution of New World dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)
Cupello M., Silva F.A.B., Vaz-de-Mello F.Z. 2023. Front. Ecol. Evol. 11: 1‒42.
2.Catalogue of Life
3.Illustrated keys to the Scarabaeinae (Coleoptera: Scarabaeidae) of Maryland
Nemes S.N., Price D.L. 2015. Northeastern Naturalist 22: 318‒344.