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For Insects, Spiders & Their Kin
For the United States & Canada
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Family Heteroceridae - Variegated Mud-loving Beetles

5014326 Heterocerid - Heterocerus fenestratus Small Beetle - Heterocerus fenestratus Small beetle with patterned elytra - Heterocerus Heterocerid - Heterocerus orange-spotted beetle - Heterocerus A Heterocerid beetle? Genus Heterocerus perhaps? - Heterocerus Beetle IMG_9650 - Heterocerus
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
Superfamily Byrrhoidea (Pill, Water and Toe-winged Beetles)
Family Heteroceridae (Variegated Mud-loving Beetles)
Synonyms and other taxonomic changes
for most recent treatment see(1)
Note: The classification proposed in(1) is reflected in the Guide. Summary of changes:
The genus Tropicus is retained in its current circumscription
Species currently treated in Augyles, Centuriatus, Explorator, and Microaugyles all go to the genus Augyles
The remaining New World species go (back!) to Heterocerus
Explanation of Names
Heteroceridae W.S. MacLeay 1825
~40 spp. in 3 genera in our area(2)(1), ~500 spp. in 15 genera worldwide(3)
sandy or muddy shores of rivers, lakes, and ponds
Print References
Pacheco F. (1964) Sistemática, filogenia y distribución de los Heteroceridos de América (Coleoptera: Heteroceridae). Monogràfías del Colegio de Postgraduados, Chapingo, Mexico. 115 pp.
Pacheco F. (1969) A new species of Heterocerini (Coleoptera: Heteroceridae). Florida Entomol. 52: 37-39. (Full text)
Miller W.V. (1988a) A new species of Heterocerus recorded from Florida and Canada (Coleoptera: Heteroceridae). Florida Entomol. 71: 30-33. (Full text)
Miller W.V. (1988b) Two new species of Heterocerus from North America, with notes on related species (Coleoptera: Heteroceridae). Col. Bull. 42: 313-320. (Full text)
Miller W.V. (1994) New species of Heteroceridae (Coleoptera) from Canada and the United States. Col. Bull. 48: 11-18. (Full text)
Schaeffer K.F., Drew W.A. (1964) The Heteroceridae (Coleoptera) of Oklahoma. Proceedings of the Oklahoma Academy of Sciences 45: 69-71. (Full text)
Works Cited
1.Molecular data resolves relationships within Heteroceridae (Coleoptera: Dryopoidea)
J.G. King, J.R. Starr, P.K. Lago. 2011. Systematic Entomology 36(3): 435–445.
2.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). 2002. CRC Press LLC, Boca Raton, FL.
3.Order Coleoptera Linnaeus, 1758. In: Zhang Z.-Q. (ed.) Animal biodiversity: An outline of higher-level classification...
Slipinski S.A., Leschen R.A.B., Lawrence J.F. 2011. Zootaxa 3148: 203–208.
4.The variegated mud-loving beetles (Heteroceridae) of Mississippi and Alabama, with discussion and keys to se US spp.
King and Lago. 2012. Insecta Mundi 0275: 1-53.
5. A catalog of the Coleoptera of America north of Mexico. Family Heteroceridae.
Pacheco, F. 1978. USDA, Agric. Handbook. x + 9 pp. 1978. USDA.