Subfamily Anomaloninae

Theriinae(1)

2 cosmopolitan tribes. 16 genera with 201 described spp. in our area; 39 genera worldwide.(2)

Small to large species. Head with clypeus convex, not separated by a groove and often coming to a median point; labrum not projecting conspicuously ventral to clypeal margin and lacking median notch; lateral ocelli near the occipital ridge. Antennae with more than 13 flagellomeres, not flattened and widened medially. Thorax with pronotum lacking bifurcate or bilobate process; propleuron with strongly produced lobe touching or overlapping pronotum; mesopleuron lacking sternaulus; postpectal carina usually complete; scutellum without median apical spine; mesosoma usually coarsely punctate; propodeum reticulated, lacking carina, and, with apex projecting between metacoxae. Fore wing 2-25 mm long with venation complete; vein 2m-cu tubular, at least in part; no closed areolet; 2/Rs usually apical to vein 2m-cu (in some species opposite or basal); 1m-cu and Rs+M hardly angled and often giving the appearance of a single vein. Metatarsal claws less than half as long as tarsomere 5. Abdomen with strong lateral compression; the first gastral segment particularly long and slender with both glymma and tergal-sternal suture absent and with the spiracle at the posterior. Ovipositor variable in length (from the height of the metasomal apex to the length of the hind tibia); dorsal subapical notch present.(1)(3)

Wing venation of Therion Wing venation of Anomalon

Worldwide(1)

Often found in drier habitats than typical for other Ichneumonidae.(1)

Koinobiont endoparasitoids of Lepidoptera and Coleoptera larvae(4)(1) Those of Anomalonini (genus Anomalon) are paraditoids of Tenebrionid beetles while members of Gravenhorstiini (all other Anomaloninae) are generalist parasitoids of Lepidoptera.(5)

The female oviposits into the host larva. The offspring emerge after the host has metamorphosed into a pupa.(1)(5)