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Family Psephenidae - Water Penny Beetles

flat larva - Psephenus herricki water penny beetle - Psephenus herricki Water Penny Beetle - Psephenus herricki Water Penny Beetle - Psephenus herricki non-gilled Water Penny Beetle - Ectopria nervosa black and orange beetle - Ectopria nervosa - female 8016534 8016534
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Longhorn, Leaf and Snout Beetles)
Superfamily Byrrhoidea (Pill, Water and Toe-winged Beetles)
Family Psephenidae (Water Penny Beetles)
Other Common Names
Long-toed Water Beetles (1)
Synonyms and other taxonomic changes
Psephenidae Lacordaire, 1854 (ITIS)
Explanation of Names
Water penny: larvae are flat and circular, and live in water
Psephenidae is from genus Psephenus Haldeman, that from Greek psepheno dark, obscure and/or psepho meaning "a small stone" or darkness (2).
Numbers
16 species in 6 genera in North America (3)
Size
typically 3-6 mm
Range
in North America, most species occur in the west
also found in several regions of the world; greatest diversity in the Orient
Habitat
Larvae are aquatic and cling to the underside of rocks in fast-flowing streams.
Adults crawl over rocks and vegetation along streams.
Food
larvae graze on algae
Print References
Arnett, American Beetles, Vol. 2, Chapter 48 (4)
Borror, entries for psepheno, psepho (2)
Ciegler, p. 177 (5)
Dillon and Dillon, p. 365, plate 18--adult of Psephenus herricki (1)
Internet References
Animal Diversity Web--Psephenidae
live larva image and distinguishing characteristics of larva (North Carolina State University)
live larva images showing upper and lower surfaces (Danielle Love, Western Washington Univ.)
Works Cited
1.A Manual of Common Beetles of Eastern North America
By Dillon, Elizabeth S., and Dillon, Lawrence
2.Dictionary of Word Roots and Combining Forms
By Donald J. Borror
3.Nearctica: Nomina Insecta Nearctica
4.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
By Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.)
5.Water Beetles of South Carolina
By Janet Ciegler