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Genus Amara - Seed-Eating Ground Beetles

Amara sp. - Amara Amara? - Amara aenea Amara sp. - Amara Black Beetle - Amara aenea Amara Ground Beetle - Amara metallic green beetle - Amara Beetle - Amara angustata - male
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Adephaga (Ground and Water Beetles)
Family Carabidae (Ground Beetles)
Subfamily Harpalinae
Supertribe Pterostichitae
Tribe Zabrini (Seed-Eating Ground Beetles)
Genus Amara (Seed-Eating Ground Beetles)
Explanation of Names
Amara Bonelli 1810
Greek a- (privative) + mairo 'to shine'(1)
Numbers
105 spp. in 11 subgenera in our area, ~600 spp. in 47 subgenera worldwide(1)
not yet in the guide: subg. Neopercosia: A. fortis LeConte
Size
5-11 mm(2), usually 6-9 mm
Identification
Medium-sized, usually oval beetles, many with coppery reflection, though some are black or brown(3). Typically encountered running about in the open. Most fully winged (macropterous)(4).
Species-level identification is difficult, even with technical keys.
Range
mostly Holarctic; across NA(4)
Habitat
Commonly found on wet sand/mud or beneath logs or rocks in damp areas(3), though tend to favor dry open areas in these habitats(4)
Food
Most feed on seeds; a few (e.g., A. obesa), on grasshopper eggs(5)(3)
Life Cycle
Some species overwinter as adults(5)
Works Cited
1.Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico
Bousquet Y. 2012. ZooKeys 245: 1–1722.
2.A Manual of Common Beetles of Eastern North America
Dillon, Elizabeth S., and Dillon, Lawrence. 1961. Row, Peterson, and Company.
3.Ground Beetles and Wrinkled Bark Beetles of South Carolina
Janet Ciegler. 2000. Clemson University.
4.American Beetles, Volume I: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia
Arnett, R.H., Jr., and M. C. Thomas. (eds.). 2000. CRC Press LLC, Boca Raton, FL.
5.An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana.
Blatchley, W. S. 1910. Indianapolis,Nature Pub. Co.