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Genus Ululodes - Split-eyed Owlflies

Owlfly - Ululodes floridanus - female Four-spotted Owlfly - Ululodes quadripunctatus - female Four-spotted Owlfly - Ululodes quadripunctatus - male Four-spotted Owlfly - Ululodes quadripunctatus - female Interesting Eyes - Ululodes macleayanus - male Owlfly eggs and hatchllings - Ululodes
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Neuroptera (Antlions, Lacewings, and Allies)
Suborder Myrmeleontiformia (Antlions and Allies)
Family Myrmeleontidae (Antlions & Owlflies)
Subfamily Ascalaphinae (Owlflies)
Genus Ululodes (Split-eyed Owlflies)
Explanation of Names
Ululodes Smith 1900
from Ulula Rambur, 1842 (an owlfly genus-group name) + Greek suffix -ειδής (-odes) 'resembling'
from the Greek ulula ('screech owl')
Numbers
7 spp. in our area(1) (and comments by J.R. Jones)
1. Ululodes arizonensis Banks, 1907: AZ
2. Ululodes bicolor (Banks, 1895): AZ, CA; records from LA, TX in older literature correspond to separate species (e.g. U. floridanus)
3. Ululodes floridanus (Banks, 1906): AL, FL, GA, LA, NC, SC, TX
4. Ululodes macleayanus (Guilding, 1825): AL, AR, FL, GA, IL, KS, LA, MO, MS, NC, NJ, OK, SC, TX
5. Ululodes mexicanus (McLachlan, 1871): AZ, CA, NM, NV, w. TX, UT
6. Ululodes nigripes Banks, 1943: TX
7. Ululodes quadripunctatus (Burmeister, 1839): widespread, especially eastern (ON; AL, AR, AZ, DC, DE, FL, IA, IL, IN, KS, KY, LA, MD, MI, MO, MS, NC, NE, NJ, NY, OH, OK, PA, SC, TN, TX, VA, WI, WV)
Identification
The eyes being divided (or "split") distinguishes this from all other owlfly genera in our area.(2)

In U. quadripunctatus, at least, apparently only male hindwings are stalked(3); female wings are not stalked.

Species identification follows unpublished revision to the genus by Oswald & Jones. Earlier keys such as that in (4) are rendered inadequate under this revision.
fore and hind pterostigmas both dark: U. arizonensis (western, wings distinctively pigmented), U. macleayanus (eastern: widespread), U. mexicanus (western, wings unpigmented), U. nigripes (TX)
fore pterostigmas pale and hind dark: U. bicolor (western), U. floridanus (eastern)
fore and hind pterostigmas both pale: U. quadripunctatus
                 
     U. arizonensis                    U. bicolor                      U. floridanus                U. macleayanus
           
     U. mexicanus                    U. nigripes                U. quadripunctatus
Range
transcontinental (WI‒ON‒MA to FL‒TX‒CA)
Remarks
Our only representative of split-eyed antlion (tribes Ululodini and Ascalaphini), that together contain ~340 spp. worldwide