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Family Aderidae - Ant-like Leaf Beetles

Antlike Leaf Beetle - Zonantes nubifer Antlike Leaf Beetle - Zonantes fasciatus Leaf Beetle  - Zonantes subfasciatus Ariotus subtropicus Casey - Ariotus subtropicus Elonus basalis Cnopus Antlike Leaf Beetle - Zonantes nubifer Aderidae? from rotting Quercus velutina wood - Ganascus ventricosus
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
No Taxon (Series Cucujiformia)
Superfamily Tenebrionoidea (Fungus, Bark, Darkling and Blister Beetles)
Family Aderidae (Ant-like Leaf Beetles)
Synonyms and other taxonomic changes
Xylophilidae Shuckard 1840, Euglenesidae (=Euglenidae) Seidlitz 1875, Hylophilidae Pic 1900
Revised in(1) but many western species still undescribed(2)
Tribal and generic limits remain unclear; world fauna has to be examined to resolve these limits(2)
Explanation of Names
Aderidae Csiki 1909
Numbers
4 tribes, with ~50 species in 11 genera in our area(2) and ~900 spp. in 50 genera total(3)
Overview of nearctic genera (* –taxa not yet in the guide)
Family Aderidae
Tribe Aderini
Subtribe Cnopina Cnopus
Subtribe Aderina Aderus
Incertae sedis "Xylophilus" constrictus

Species checklist
Tribe Phytobaenini Báguena 1948
Genus Axylophilus Casey 1895
A. yuccae Casey 1895
Genus Ganascus Casey 1895
G. ventricosus (LeConte 1875) – Type
G. ptinoides (Schwarz 1878) – Type
Tribe Euglesinini Seidlitz 1875
Genus Zonantes Casey 1895
Z. subfasciatus (LeConte 1855)
Z. pallidus Werner 1990
Z. floridanus Werner 1990
Z. signatus (Haldeman 1848) – Type
Z. nubifer (LeConte 1878) – Type
Z. fasciatus (Melsheimer 1846)
Z. hubbardi Casey 1895
Z. ouachitanus Werner 1990
Z. ater (LeConte 1875) – Type
Z. mississippiensis Werner 1990
Tribe Emelinini Báguena 1948
Genus Emelinus Casey 1895
E. butleri Werner 1956 – Type
E. huachucanus Werner 1956 – Type
E. melsheimeri (LeConte 1855) – Type
Tribe Aderini Winkler 1927
Subtribe Cnopina Báguena 1948
Genus Cnopus Champion 1893
C. impressus (LeConte 1875) – Type
C. nucleus (Fall 1901) – Type
Subtribe Aderina Winkler 1927
Genus Aderus Westwood 1830
A. brunnipennis (LeConte 1875)
A. populneus (Creutzer 1796)
A. saginatus (Casey 1895)
A. tantillus (Champion 1890)
Subtribe Syzetoninina Báguena 1948
Genus Vanonus Casey 1895
V. huronicus Casey 1895
V. brunnescens Fall 1901
V. macrops Werner 1990
V. musculus Werner 1990
V. oklahomensis Werner 1990
V. sagax Casey 1895
V. valgus Werner 1990
V. vigilans Casey 1895
V. wickhami Casey 1895
V. calvescens Casey 1895
V. piceus (LeConte 1855) – Type
V. uniformis Werner 1990
V. balteatus Werner 1990
Genus Ariotus Casey 1895
A. luteolus (Casey 1895)
A. pruinosus (Casey 1895)
A. quercicola (Schwarz 1878)
A. subtropicus Casey 1895
Genus Pseudariotus Casey 1895
P. notatus (LeConte 1855) – Type
Genus Elonus Casey 1895
E. basalis (LeConte 1855) – Type
E. chisosensis Werner 1992
E. excavatus Werner 1992
E. hesperus Werner 1990
E. nebulosus (LeConte 1875) – Type
E. simplex Werner 1992
Subtribe Gompeliina Bouchard 2011 (=Olotelina Báguena 1948)
Genus Gymnoganascus Werner 1990
G. stephani Werner 1990
Incertae sedis
Size
1-4 mm(2)
Identification
Slightly ant-like, oval to elongate. Head strongly deflexed, wider than long, and wider than pronotum, abruptly constricted at base in most. Eyes large and coarsely granulate and weakly to strongly emarginate (margin indented) in most with setae (hairs) between facets. Basal two sternites (abdominal plates) are fused.
Range
Worldwide.
Habitat
Adults can often be found on the underside of leaves of various trees and shrubs, and by beating and sweeping vegetation. Several species seem to be attracted to oaks and at least one has been found on Yucca. Larvae in rotting wood, bee nests, termite nests, leaf litter, under bark.(2)

Adults are also often collected using Malaise and flight-intercept traps, and by using pyrethin knockdown sampling. (4)
Life Cycle
Larvae poorly known. Only those of 3 genera have been thoroughly described. (2)

One genus (Megaxenus, from Australia & Papua New Guinea) is termitophilus. Larvae of Vanonus balteatus were found in the nesting substrate of a bee nest in Mexico, probably feeding on fungi associated with decaying nesting material. (4)
Internet References
(5)
Works Cited
1.Revision of the Aderidae of eastern North America.
Werner, F.G. 1990. Journal of the New York Entomological Society, 98: 187-232.
2.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). 2002. CRC Press LLC, Boca Raton, FL.
3.Order Coleoptera Linnaeus, 1758. In: Zhang Z.-Q. (ed.) Animal biodiversity: An outline of higher-level classification...
Ślipiński S.A., Leschen R.A.B., Lawrence J.F. 2011. Zootaxa 3148: 203–208.
4.Coleoptera, Beetles; Morphology and Systematics vol. 2
Richard A.B. Leschen, Rolf G. Beutel and John F. Lawrence. 2010. De Gruyter.
5.Gompel N. (2010) An iconography of Aderidae species from North America and Western Europe