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Subfamily Doryctinae

Pretty Wings - Spathius - female braconid - female Braconidae - Spathius - female Female braconid? - female Braconid wasp - Doryctinae - female Braconid - Spathius - male Spathius elegans - female Spathius elegans? - Spathius - female
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hymenoptera (Ants, Bees, Wasps and Sawflies)
No Taxon ("Parasitica" - Parasitoid Wasps)
Superfamily Ichneumonoidea (Braconid and Ichneumonid Wasps)
Family Braconidae (Braconid Wasps)
Subfamily Doryctinae
Synonyms and other taxonomic changes
Largely unrevised since the Krombein et al. (1979) Catalog. A single new species included in Nomina Insecta Nearctica (1996): Heterospilus frommeri Marsh, 1989.

Histeromerus Wesmael, 1838 (1 sp.) has been removed from Doryctinae to the monotypic Histeromerinae.
Acrophasmus Enderlein, 1912 (7 spp.) is treated as a junior synonym of Doryctinus.
Aptenobracon (1 sp.) is treated as a junior synonym of Rhaconotus.
Numbers
159 spp. in 25 genera in our area;(1)(2) more than 2000 spp. in 198 genera and 15 tribes worldwide
Subfamily Doryctinae
Tribe Binareini
1. Liobracon (1 sp.)

Tribe Doryctini
Subtribe Dendrosotina
2. Dendrosoter (6 spp.)
Subtribe Doryctina
3. Doryctes (15 spp.)
4. Ontsira (8 spp.)
5. Pedinotus (1 sp.)
6. Rhoptrocentrus (1 sp.)

Tribe Ecphylini
7. Aivalykus (1 sp.)
8. Ecphylus (22 spp.)

Tribe Hecabolini
Subtribe Hecabolina
9. Hecabolus (1 sp.)
10. Leluthia (3 spp.)
11. Monolexis (1 sp.)
Subtribe Pambolideina
12. Pambolidea (2 spp.)
Subtribe Stenocorsina
13. Allorhogas (4 spp.)
14. Callihormius (4 spp.)
15. Curtisella (1 sp.)
16. Doryctinus (9 spp.)
17. Glyptocolastes (2 spp.)
18. Stenocorse (1 sp.)

Tribe Heterospilini
19. Heterospilus (32 spp.)
20. Pioscelus (2 spp.)

Tribe Holcobraconini
Subtribe Odontobraconina
21. Odontobracon (6 spp.)

Tribe Rhaconotini
Subtribe Rhaconotina
22. Aptenobracon (1 sp.)
23. Rhaconotus (9 spp.)

Tribe Spathiini
Subtribe Ptesimogastrina
24. Ptesimogaster (2 spp.)
Subtribe Spathiini
25. Spathius (24 spp.)
Size
1-25 mm (not including ovipositor)(3)
Identification
Row of thickened spines (figures 9, 10) along length of outer or front surface of fore tibia and sometimes also middle tibia (these may be a challenge to find)(Belokobilskij and Zaldivar-Riveron 2014)
"Cyclostome mouth": short clypeus + concave labrum above mandibles resembles a found, open mouth. See Remarks on Braconidae page.
It is one of the most diverse and species-rich subfamilies of the Braconidae. The subfamily is diagnosed by a row of spines on the fore tibia, a flange on the propleuron above the forecoxa, the presence of an epicnemial carina and occipital carina (Marsh, 2002), a double node near the apex of the dorsal valve of the ovipositor (Quicke et al., 1992), and a cyclostome mouth. (Seltmann, K. and M.J. Sharkey, 2007).
Food
Known larval hosts: larvae of beetles, sawflies, and some moths.(3)
Life Cycle
Largely unknown as a subfamily. Those that are known are idiobiont ectoparasitoids.(3)
Print References
Belokobylskij, S.A. and A. Zaldivar-Riveron. 2014. The genus Spathius Nees (Hymenoptera, Braconidae, Doryctinae) in Mexico: occurrence of a highly diverse Old World taxon in the Neotropics. Zookeys 427: 59-73. (Full text)
Edmardash, Y.A.; El-Ghiet, U.M.A.; Soliman, A.M.; Al-Fifi, Z.I.A.; & Gadallah, N.G. (2020). First contribution to the doryctine fauna (Hymenoptera, Braconidae, Doryctinae) of Farasan Archipelago, Saudi Arabia, with new records and the description of a new species. ZooKeys, 977: 41–74.
Johnson, N. F., & Triplehorn, C. A. (2005). Borror and DeLong's Introduction to the Study of Insects.(4)
Marsh, P. M. (1965). The Nearctic Doryctinae. I. A Review of the Subfamily with a Taxonomic Revision of the Tribe Hecabolini (Hymenoptera: Braconidae). Annals of the Entomological Society of America 58(5): 668–699.
Seltmann, K. and M.J. Sharkey. 2007. A new genus and species of apterous Doryctinae (Hymenoptera: Braconinae) from Costa Rica. Zootaxa 1415: 17-24. (Full text)
Wharton, R., Marsh, P., & Sharkey, M. (1997). Identification manual to the New World genera of the family Braconidae (Hymenoptera). International Society of Hymenopterists Special Publication.(3)
Works Cited
1.Family Braconidae, pp. 144-295. In: Catalog of Hymenoptera in America North of Mexico
Marsh, P.M. . 1979. Smithsonian Institution Press.
2.Nomina Insecta Nearctica
Poole, Robert W. Nearctica.com, Inc.
3.Manual of the New World Genera of the Family Braconidae (Hymenoptera)
Wharton, R.A., P.M. Marsh, M.J. Sharkey (Eds). 1997. International Society of Hymenopterists.
4.Borror and DeLong's Introduction to the Study of Insects
Norman F. Johnson, Charles A. Triplehorn. 2004. Brooks Cole.