Includes Aphidoletini, Asphondyliini, Cecidomyiini, Clinodiplosini, Lestodiplosini, Mycodiplosini, and several unplaced genera.
Typically 12 flagellomeres. Male flagellomeres are binodal (except in most Asphondyliini) and their circumfila are looped. Females have cylindrical flagellomeres. Most adults have a long, narrow lobe at the top of the head behind the eyes, which bears a pair of large proclinate setae. Female cerci are free, but may be closely appressed or so reduced that they appear to be fused. The aedeagus is free and appears more sclerotized than in the Lasiopteridi. Ventral papillae of the larvae are generally asetose. (1)
This group contains gall makers, fungus feeders, and predators. (1)