Includes the tribes Alycaulini, Lasiopterini, Ledomyiini, Oligotrophini, and Trotteriini.
Most have an irregular number of flagellomeres, rarely numbering exactly 12. Male flagellomeres are made up of a node and a distal neck that is longer than on female antennae. The cerci of the ovipositor are usually fused into one caudal lobe. The male aedeagus is supported on each side by gonocoxal lobes. Most ventral papillae of the larvae are setose. (1)
Mostly gall makers; a few fungus feeders.