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Order Psocodea - Barklice, Booklice, and Parasitic Lice

Bark Louse with Yellow Eyes - Xanthocaecilius quillayute Yellow-eyed barklouse - Xanthocaecilius Barklouse - Aaroniella badonneli Immatures - Ectopsocus californicus possible booklouse - Camelopsocus tucsonensis - female False Lizard Barklouse - Pseudocaecilius citricola Brachypterous Barklouse 2 - Psyllipsocus ramburii Barklice - Lachesilla andra - male
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Psocodea (Barklice, Booklice, and Parasitic Lice)
Synonyms and other taxonomic changes
=Psocoptera, Copeognatha, Corrodentia
Note: Information on this page doesn not fully reflect the incorporation of the parasitic lice, Phthiraptera, formerly often treated as two separate orders, Mallophaga (chewing lice) and Anoplura (sucking lice). See Tree of Life for the current concept of relationships among these groups. However, Phthiraptera and Psocoptera are still sometimes treated by leading experts as two orders that together form the superorder Psocodea(1).
Explanation of Names
Greek psokos, "rubbed small, gnawed" (perhaps referring to manner of eating) plus pteron, wing(2)
Numbers
3 suborders, with ~1,220 spp. in >200 genera of 43 families in our area (incl. ~920 spp., >130 genera and 17 families of parasitic lice)(2)(3) and ~8,000 spp. in TBA genera of ~80 families worldwide
Overview of our fauna
Taxa not yet in the guide: * native, + non-native. Classification adapted from(4)
Order Psocodea
Suborder TROGIOMORPHA
Infraorder ATROPETAE
Subfamily THYLACELLINAE Thylacella Enderlein 1911

Subfamily *PERIENTOMINAE *Proentomum Badonnel 1949, *Soa Enderlein 1904
Subfamily LEPIDOPSOCINAE Echmepteryx Aaron 1886

Subfamily ECHINOPSOCINAE Neolepolepis Mockford 1993, Pteroxanium Enderlein 1922

Family TROGIIDAE Cerobasis Kolbe 1882, Lepinotus Heyden 1850, *Myrmicodipnella Enderlein 1909, Trogium Illiger 1798

Family PSOQUILLIDAE +Balliella Badonnel 1949, +Psoquilla Hagen 1865, Rhyopsocus Hagen 1876

Infraorder PSOCATHROPETAE
Family PSYLLIPSOCIDAE Dorypteryx Aaron 1883, Pseudorypteryx Garcia Aldrete 1984, +Psocathropos Ribaga 1899, Psyllipsocus Selys-Longchamps 1872

Family PRIONOGLARIDIDAE Speleketor Gurney 1943

Suborder TROCTOMORPHA
Infraorder AMPHIENTOMETAE
Subfamily AMPHIENTOMINAE Lithoseopsis Mockford 1993, Stimulopalpus Enderlein 1906

Infraorder NANOPSOCETAE
Subinfraordinal group PHTHIRAPTERA... TBA
Superfamily group Amblycera
Superfamily group Ischnocera
Superfamily group Anoplura

Subfamily EMBIDOPSOCINAE Belaphotroctes Roesler 1943, Embidopsocus Hagen 1866

Subfamily LIPOSCELIDINAE Liposcelis Motschulsky 1852

Subfamily +PACHYTROCTINAE +Pachytroctes Enderlein 1905
Subfamily TAPINELLINAE Nanopsocus Pearman 1928, Tapinella Enderlein 1908

Family SPHAEROPSOCIDAE +Badonnelia Pearman 1953, *Prosphaeropsocus Mockford 2009, Sphaeropsocus Hagen 1882, *Troglosphaeropsocus Mockford 2009

Suborder PSOCOMORPHA
Infraorder EPIPSOCETAE
Family EPIPSOCIDAE Bertkauia Kolbe 1882, Epipsocus Hagen 1866

Infraorder CAECILIUSETAE
Superfamily Asiopsocoidea
Family ASIOPSOCIDAE Asiopsocus Günther 1968

Superfamily Caeciliusoidea
Subfamily AMPHIPSOCINAE Polypsocus Hagen 1866

Family DASYDEMELLIDAE Teliapsocus Chapman 1930

Subfamily CAECILIUSINAE Tribe +Epicaeciliini: +Orocaecilius Mockford 2000; Tribe Coryphacini: Stenocaecilius Mockford 2000, Valenzuela Navás 1924; Tribe Maoripsocini: Maoripsocus Tillyard 1923

Subfamily PARACAECILIINAE Xanthocaecilius Mockford 1989

Family STENOPSOCIDAE Graphopsocus Kolbe 1880

Infraorder HOMILOPSOCIDEA
Subfamily EOLACHESILLINAE Anomopsocus Roesler 1940, *Nanolachesilla Mockford and Sullivan 1986, *Prolachesilla Mockford and Sullivan 1986

Subfamily LACHESILLINAE Lachesilla Westwood 1840

Family ECTOPSOCIDAE Ectopsocopsis Badonnel 1955, Ectopsocus McLachlan 1899

Family PERIPSOCIDAE *Kaestneriella Roesler 1943, Peripsocus Hagen 1866

Family TRICHOPSOCIDAE Trichopsocus Kolbe 1882

Subfamily ARCHIPSOCINAE *Archipsocopsis Badonell 1948, Archipsocus Hagen 1882

Subfamily PSEUDOCAECILIINAE Ophiodopelma Enderlein 1908, Pseudocaecilius Enderlein 1903

Subfamily AARONIELLINAE Aaroniella Mockford 1951

Subfamily PHILOTARSINAE Philotarsus Kolbe 1880

Family ELIPSOCIDAE
Subfamily ELIPSOCINAE Cuneopalpus Badonnel 1943, Elipsocus Hagen 1866

Subfamily *NEPIOMORPHINAE *Nepiomorpha Pearman 1936
Subfamily PROPSOCINAE Propsocus McLachlan 1866

Subfamily PSEUDOPSOCINAE *Palmicola Mockford 1955, Reuterella Enderlein 1903

Family MESOPSOCIDAE Mesopsocus Kolbe 1880

Infraorder PSOCETAE
Family HEMIPSOCIDAE Hemipsocus Selys-Longchamps 1872

Family MYOPSOCIDAE Lichenomima Enderlein 1910, Myopsocus Hagen 1866

Family PSOCIDAE
Subfamily AMPHIGERONTIINAE Amphigerontia Kolbe 1880, Blaste Kolbe 1883, Blastopsocus Roesler 1943

Subfamily PSOCINAE Tribe Cerastipsocini: Cerastipsocus Kolbe 1884; Tribe Metylophorini: Metylophorus Pearman 1932; Tribe Psocini: *Atropsocus Mockford 1993, Hyalopsocus Roesler 1954, Psocus Latreille 1794; Tribe Ptyctini: Camelopsocus Mockford 1965, Indiopsocus Mockford 1974, Loensia Enderlein 1924, Ptycta Enderlein 1925, Steleops Enderlein 1910, Trichadenotecnum Enderlein 1909
Size
typically 1.5-5 mm (up to 10 mm; rarely under 1 mm)
Identification
Good profiles of Psocoptera and Phthiraptera in(5)
Keys to NA bark- and booklice in(6)
Keys to families in(7)(2)
Pictorial guide to TX fauna in(8)
A helpful interactive key to adults in(9) (covers the fauna of the UK and Ireland)
Pictorial key to pest booklice in(10)
Images of many cosmopolitan spp. and those otherwise adventive in NA(9)
Range
worldwide; a number of species cosmopolitan and spreading through the agency of man
Habitat
barklice are always found outdoors, and occupy a wide variety of habitats; booklice often live in homes (more here)
Food
Barklice feed on lichens and fungi on tree bark, booklice are best known for feeding on the starch in book bindings(7)
Life Cycle
metamorphosis gradual (paurometabolous insects)(7)