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Family Geotrupidae - Earth-Boring Scarab Beetles

Geotrupes blackburnii Odonteus obesus - female Bolbelasmus minor (Linell) - Bolbelasmus minor Shiny Digging Beetle - Geotrupes blackburnii ui scarab - Eucanthus Ceratophyus gopherinus Cartwright - Ceratophyus gopherinus - male Geotrupes - Geotrupes blackburnii Earth-borer - Bolbocerosoma
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
Superfamily Scarabaeoidea (Scarab, Stag and Bess Beetles)
Family Geotrupidae (Earth-Boring Scarab Beetles)
Synonyms and other taxonomic changes
Until recently treated as a subfamily of Scarabaeidae
Explanation of Names
Geotrupidae Latreille 1802
Numbers
55 spp. in 11 genera in the Nearctic(1), ~620 spp. in ~70 genera worldwide(2) arranged into 13 tribes and 3 subfamilies(3)
Overview of our fauna:
Family GEOTRUPIDAE
Subfamily BOLBOCERATINAE
Odonteus 10 spp., mostly eastern
Eucanthus 5 spp., mostly eastern
Bradycinetulus 3 spp., eastern/central US
Bolbocerastes 3 spp., w. US
Bolbocerosoma 10 spp., eastern/central US
Bolbelasmus 4 spp., sw. US
Bolborhombus 3 spp., sw. US
Subfamily GEOTRUPINAE
Geotrupes 9 spp., US except far west
Mycotrupes 5 spp., se. US
Peltotrupes 2 spp., Florida
Ceratophyus 1 sp., California
Size
5-45 mm(2)
Identification
Eleven-segmented antennae, unlike the Scarabs which have fewer segments.

Bodies are stout and often hemispherical.
Subfamilies:
Geotrupinae (left): Antennal club small, body largely black with or without metallic sheen
Bolboceratinae (right): Antennal club large, body brown, some with black markings
Habitat
These beetles spend most of their lives in burrows one to four feet down, often under dung or carrion.
Food
Larvae feed on dung or carrion. Adults feed on dung or fungi, or do not feed at all.
See Also
Internet References
Family overview(2) - broken link
Works Cited
1.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). 2002. CRC Press LLC, Boca Raton, FL.
2.Generic Guide to New World Scarab Beetles (by Brett Ratcliffe and Mary Liz Jameson)
3.Family-group names in Coleoptera (Insecta)
Bouchard P. et al. 2011. ZooKeys 88: 1–972.