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Family Geotrupidae - Earth-Boring Dung Beetles

Golden Scarab - Geotrupes splendidus Dung Beetle - Geotrupes balyi Geotrupes hornii Which Geotrupes? - Geotrupes splendidus could it be? - Family Ochodaeidae - Sand-loving Scarab Beetles [30] - Odonteus - female Beetle - Eucanthus School project #1 - Geotrupes splendidus BG2578 E2978 - Geotrupes blackburnii
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Longhorn, Leaf and Snout Beetles)
Superfamily Scarabaeoidea (Scarab, Stag and Bess Beetles)
Family Geotrupidae (Earth-Boring Dung Beetles)
Synonyms and other taxonomic changes
Until recently these beetles were included in the Scarabaeidae family, as subfamily Geotrupinae.
Numbers
11 Nearctic genera, 55 Nearctic species.

Family GEOTRUPIDAE
Subfamily BOLBOCERATINAE
Odonteus - 10 species, generally distributed, most in the east
Eucanthus - 5 species, generally distributed, most in the east
Bradycinetulus - 3 species, eastern US, to Nebraska and Texas
Bolbocerastes - 3 species, western US
Bolbocerosoma - 10 species, western US
Bolbelasmus - x species, western US
Bolborhombus - x species, western US
Subfamily GEOTRUPINAE
Geotrupes - 9 species, US, except far western states
Mycotrupes - 5 species, SE US
Peltotrupes - 2 species, Florida endemics
Ceratophyus - 1 species, C. gopherinus, California endemic
Size
Typically 10-17 mm.
Identification
Eleven-segmented antennae, unlike the Scarabs which have fewer segments.

Bodies are stout and often hemispherical.

Two subfamilies, as follows
Geotrupinae (left): Antenna with club small. Body largely black with or without metallic sheen.
Bolboceratinae (right): Antenna with club large. Body brown, some with black markings.
Habitat
These beetles spend most of their lives in burrows one to four feet down, often under dung or carrion.
Food
Larvae feed on dung or carrion. Adults feed on dung or fungi, or do not feed at all.
Print References
Peterson Field Guide (1)
American Beetles, Vol. 2, Chapter 29 (2)
Howden, H.F. 1955. The biology and taxonomy of the North American beetles of the subfamily Geotrupinae with revisions of the genera Bolbocerosoma, Eucanthus, Geotrupes, and Peltotrupes (Scarabaeidae). Proceedings of the United States National Museum 104: 151-319.
Works Cited
1.Peterson Field Guides: Beetles
By Richard E. White
2.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
By Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.)