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Genus Chlaenius - Vivid Metallic Ground Beetles

Vivid Metallic Ground Beetles - Chlaenius aestivus - male - female Velvety green carabid Chlaenius sericeus - Chlaenius sericeus Ground beetle - Chlaenius emarginatus C. tricolor? - Chlaenius tricolor Chlaenius alternatus Green Metallic ground beetle - Chlaenius sericeus Chlaenius - which one? - Chlaenius tricolor - male Ground beetle - Chlaenius erythropus - male
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Adephaga (Ground and Water Beetles)
Family Carabidae (Ground Beetles)
Subfamily Harpalinae
Tribe Chlaeniini
Genus Chlaenius (Vivid Metallic Ground Beetles)
Synonyms and other taxonomic changes
Chlaenius Bonelli, 1810
Explanation of Names
From Greek chlaen cloak (Word Roots and Combining Forms). This likely refers to the fine pubescence of the dorsal surface.
Numbers
Nearctica (1) lists 51 North American species.
Arnett, p. 391, lists 50 species (2).
Size
8-23 mm
Identification
Medium sized to large ground beetles. Attractive, often with vivid metallic coloration of head, pronotum. Pronotum has distinctive "punctures" (indentations). Head, pronotum, and elytra have fine hairs (setae), thus are called "setose" in technical language. Front tarsomeres 1-3 of male are dilated, have spongy underside, presumably used in mating (3) (4).
They are covered with hairs that lie flat.(5)

Some images showing these characteristics:


Reported to smell foul when disturbed (like leather), therefore probably have strong chemical defenses, like many in this family.
Range
Much of North America? Perhaps 20 species in east.
Habitat
Usually moist areas, some semiaquatic.
Season
Typically spring-early summer, again in late summer-fall.
Food
Predatory.
Life Cycle
Some species lay eggs in mud cells place on underside of leaves or twigs. Cell morphology can be distinctive for a species. Presumably most species have one generation per year, with adults overwintering to breed in spring. Come to lights.
Print References
Arnett, p. 391, descr. 2 species, fig. 24.38--C. pensylvanicus (2)
Ciegler, pp. 78-82, describes and keys 18 spp. (3)
Dillon, p. 109 (key to 9 spp.), plate XII, XIII--Anomoglossus (now in Chlaenius?) (4)
Arnett, et al., pp. 99-100, figs. 255-258 (6)
White, p. 92, fig. 30, lists circa 40 spp. (7)
Brimley, p. 126 (8)
Swan and Papp, pp. 344-345, figs. 568-570 (9)
Papp, p. 53, figs. 148-151 (10)
Internet References
Ground Beetles of Canada illustrates several species--these are the best photographs