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BugGuide Gathering
Pack Forest
Washington State
July 10-12, 2009
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Family Ochodaeidae - Sand-loving Scarab Beetles

Sand-loving Scarab? - Parochodaeus pectoralis blond Sand-loving Scarab? Sand-loving Scarabs + one Xenochodaeus musculus Parochodaeus duplex Uncommon ochodaeid - Codocera gnatho - male ochodaeid ochodaeid
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Longhorn, Leaf and Snout Beetles)
Superfamily Scarabaeoidea (Scarab, Stag and Bess Beetles)
Family Ochodaeidae (Sand-loving Scarab Beetles)
Pronunciation
O-ko-DEE-id-ee
Numbers
After Paulsen (2007)

Family OCHODAEIDAE Mulsant and Rey, 1871:236
Subfamily OCHODAEINAE Mulsant and Rey, 1871:236
Genus Codocera Eschscholtz, 1821:398
      =Stomphax Fischer von Waldheim 1823:158
Codocera gnatho (Fall in Fall and Cockerell, 1907:247) (Ochodaeus)
      =Ochodaeus nimius Fall, 1907:248

Genus Cucochodaeus Paulsen, 2007:9
Cucochodaeus sparsus (LeConte, 1868:51) (Ochodaeus)
      =Ochodaeus mandibularis Linell, 1896:723

Genus Neochodaeus Nikolajev, 1995:77
Neochodaeus frontalis (LeConte, 1863:76) (Ochodaeus)
      =Ochodaeus complex LeConte, 1868:51
Neochodaeus praesidii (Bates, 1887:106) (Ochodaeus)
Neochodaeus repandus (Fall, 1909:36) (Ochodaeus)
Neochodaeus striatus (LeConte, 1854:222) (Ochodaeus)

Genus Parochodaeus Nikolajev, 1995:77
Parochodaeus biarmatus (LeConte, 1868:51) (Ochodaeus)
Parochodaeus californicus (Horn, 1895:224) (Ochodaeus) type
Parochodaeus howdeni (Carlson, 1975:60) (Ochodaeus)
Parochodaeus inarmatus (Schaeffer, 1906:270) (Ochodaeus)
Parochodaeus duplex (LeConte, 1868:51) (Ochodaeus)
      =Ochodaeus kansanus Fall, 1909:34
Parochodaeus pectoralis (LeConte, 1868:51) (Ochodaeus)
Parochodaeus peninsularis (Horn, 1895:224) (Ochodaeus)
Parochodaeus ritcheri (Carlson, 1975:62) (Ochodaeus)

Genus Xenochodaeus Paulsen, 2007:3
Xenochodaeus americanus (Westwood, 1852:66) (Ochodaeus)
      =Ochodaeus opacus LeConte, 1868:51
Xenochodaeus luscinus (Howden, 1968:1118) (Ochodaeus)
Xenochodaeus musculus (Say, 1835:178) (Odontaeus)
Xenochodaeus planifrons (Schaeffer, 1906:269) (Ochodaeus)
Xenochodaeus simplex (LeConte, 1854:222) (Ochodaeus)
Xenochodaeus ulkei (Horn, 1876:182) (Ochodaeus)

Subfamily CHAETOCANTHINAE Scholtz, 1988:231
Genus Pseudochodaeus Carlson and Ritcher, 1974:99
Pseudochodaeus estriatus (Schaeffer, 1906:271) (Ochodaeus)
Identification
Ochodaeid spp. have prominent mandibles that extend past the clypeus and are visible from above; mesotibia with long spur serrate along one edge (see below):


Superficially resembles female Odonteus spp., but in addition to the above, ochodaeids won't have the divided eyes.
Range
Ochodaeidae is worldwide in temperate and tropical habitats, except Australia.
Habitat
One southeastern species is in sandy, wooded habitats. Another eastern species makes burrows in hard packed clay in wooded areas and can be found on hiking trails. Western species generally in sandy habitats. Species are generally attracted to light.
Food
Unknown. Larvae of most species have never been found.
Print References
American Beetles, Vol. 2, Chapter 30 (1)

Paulsen, M. J. 2007. Nomenclatural changes in the Nearctic Ochodaeinae and description of two new genera (Coleoptera: Scarabaeoidea: Ochodaeidae) Insecta Mundi 0021: 1-13. Download
Works Cited
1.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
By Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.)