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Family Scenopinidae - Window Flies

Window Fly - Scenopinus - male Window Fly - Scenopinus fly 43 - Belosta - female Fly - Scenopinus window fly Scenopinus - female Metatrichia bulbosa Metatrichia bulbosa
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Diptera (Flies)
No Taxon (Orthorrhapha)
Superfamily Asiloidea
Family Scenopinidae (Window Flies)
Synonyms and other taxonomic changes
Includes several genera formerly placed in Bomblyiidae.
Explanation of Names
Scenopinidae Burmeister 1835
Common name: adults of Scenopinus fenestralis are commonly found at windows inside buildings
3 subfamilies, with ~160 spp. in 9 genera in our area (>130 spp. are in 3 major genera: Brevitrichia, Pseudatrichia, & Scenopinus) and >420 spp. in 25 genera worldwide (of which almost 200 spp. are in Scenopinus)(1)
Rarely >5 mm(1)
Adult: small and dark or blackish; antennae 3-segmented, not aristate; mouth functional, non-piercing, with very short proboscis; wings with large discal cell, subapical cell, and relatively long closed anal cell; lower calypter much reduced or absent; feet without triple pad; antennae with distinctive sunken sensillae

Key to genera in(1)
Key to genera/species in (2) (does not include all species)
Various habitats; greatest diversity in arid regions with dry sandy soil, larvae in friable soils and leaf litter (have also been reared from, or suspected of breeding in, galleries of wood-boring insects, birds/mammal nests, termite mounds, bat guano, beehives and stored products). Adults of some species (e.g. Scenopinus spp.) often on windows inside homes (as larvae, they prey on pests associated with human activities)(1)
In NA, usually May-Sep
Larvae prey on small arthropods; adults take nectar from flowers, and also feed on honeydew; some adults apparently do not feed(1)
Print References
Yeates D.K. (1992) Towards a monophyletic Bombyliidae (Diptera): the removal of the Proratinae (Diptera, Scenopinidae). American Museum novitates no. 3051 (online via AMNH)
Works Cited
1.Window flies of the world (Diptera: Scenopinidae), by S. Winterton & S. Gaimari
2.A revision of the Scenopinidae (Diptera) of the world
Kelsey, L.P. 1969. Bulletin of the United States National Museum 277: 1-336.