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Family Conopidae - Thick-headed Flies

Wasp-like fly - Physocephala Fly from Adirondacks Thickheaded Fly - Physocephala Rare Conopid in California - Dalmannia vitiosa - female Physocephala sp. fly? - Physoconops obscuripennis Thick-headed Fly Conopidae - Physocephala Myopa? - Myopa
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Diptera (Flies)
No Taxon (Acalyptratae)
Superfamily Sciomyzoidea
Family Conopidae (Thick-headed Flies)
Pronunciation
kuh-nop'-uh-dee
Explanation of Names
Greek conops (κωνωψ) 'gnat'(1)(2), from conos (κωνος) 'cone' + ops (ωψ) 'eye, face'(2)
Numbers
67 spp. in 9 genera in our area(3), over 800 spp. worldwide(4)(5)
Size
4-18 mm(3)
Identification
Medium-sized, blackish or brownish flies, many of which mimic thread-waisted wasps (Sphecidae)(6)(7):
in some genera the abdomen long, with a slender, wasp-like pedicel; in others abdomen more uniform in width
head slightly broader than thorax; head often with prominent grooves on front
antennae usually long, about the same length as head and thorax combined; project forward, 3-segmented, third segment bearing either a dorsal arista or a terminal style
ocelli present or absent
proboscis long and slender, elbowed and projecting upward in some species
wing venation similar to that in the Syrphidae, but no spurious vein
Conopids may be distinguished from syrphids that lack a spurious vein by their long, slender proboscis(6).
Examples of wing venation:
  
Habitat
The adults are usually found on flowers.
Food
Adults take nectar. Larvae are endoparasites of wasps, bees, ants, crickets, cockroaches, and some Diptera (mostly calyptrate); host group varies by subfamily(5)
Life Cycle
At least in some groups, female conopids oviposit on hosts during flight. Females have complex abdominal structures to lay eggs on adult hosts prying open the exoskeleton (below left). In Stylogaster (below right), the ovipositor is long and robust, the eggs having a barbed tip for penetration and attachment of the host in the manner of a harpoon.
See Also
Print References
Camras & Hurd, "The Conopid Flies of California" (1957)
Internet References