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Family Coreidae - Leaffooted Bugs
Classification Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Class Insecta (Insects)
Order Hemiptera (True Bugs, Cicadas, Hoppers, Aphids and Allies)
Suborder Heteroptera (True Bugs)
Family Coreidae (Leaffooted Bugs)
Other Common Names Squash Bugs, Coreids
Explanation of Names Author of family is Leach, 1815. From name of genus Coreus, that New Latin (Fabricius, 1803) from Greek, koris (κορισ), a bedbug (1).
Numbers Arnett lists 33 genera and 88 species for North America (2).
Slater lists 33 genera from the United States (3).
Size Length 7-40 mm (worldwide), typically 10-20 mm, some North American species to 30 mm.
Identification Medium to large, often dark-colored hemiptera (heteroptera). Similar to Lygaeidae but with many parallel veins in membrane of front wings. Alydidae have similar venation on wings, but head nearly as wide and long as pronotum--head of Coreidae is narrower than pronotum. Family characteristics (from print and Internet references):
beak, 4-segmented
front wing with many veins, visible when folded against abdomen
Abdominal margin in many species raised--the folded wings lying in the depression formed by the margin (2).
Scent glands present on the thorax between middle and hind coxae.
Head narrower than and often shorter than the pronotum.
Hind tibiae of some species expanded and resembling leaves, e.g., Acanthocephala, Leptoglossus species. In some, e.g., Chariesterus, antennal segments are similarly expanded.
Some images showing anatomic details of the family in Leptoglossus oppositus:
Typical body forms:
Range Widely distributed but most diverse in the South and Southwest.
Food Most suck the juices of plants, but a few are reported to be predaceous.
Life Cycle Eggs are typically cemented beneath foliage of host. One or more generations per year. Adults sometimes overwinter.
Remarks Coreids often give off an unpleasant odor when handled. This is due to the secretion of defensive compounds from the thoracic glands. The active compounds of the secretions are mostly "straight-chain aldehydes and ketones--carbonyl compounds--of which trans-2-hexenal is perhaps one of the most common" (4). The defensive secretions are quite similar to those of the stink bugs, Pentatomidae (4).
A few species of Coreidae are agricultural pests.
Family COREIDAE Leach 1815
Subfamily Coreinae Leach 1815
Tribe Acanthocephalini Stål 1870
Genera Acanthocephala Laporte 1833
Tribe Acanthocerini Bergroth 1913
Genera Acanthocerus Palisot 1818, Euthochtha Mayr 1865, Sagotylus Mayr 1865
Images of Acanthocerus lobatus Burmeister 1835 (Crinocerus) (not yet in the guide) see Image #7
Description of Sagotylus confluentus Say 1832 (Coreus) (not yet in the guide) see here
Tribe Anisoscelidini Laporte 1832
Genera Chondrocera Laporte 1832, Leptoglossus Guérin-Méneville 1831, Narnia Stål 1862
Tribe Chariesterini Stål 1867
Genera Chariesterus Laporte 1833
Tribe Chelinideini Blatchley 1926
Genera Chelinidea Uhler 1863
Tribe Coreini Leach 1815
Genera Althos Kirkaldy 1904, Anasa Amyot and Serville 1843, Catorhintha Stål 1859, Centrocoris Kolenati 1845, Cimolus Stål 1862, Ficana Stål 1862, Hypselonotus Hahn 1833, Madura Stål 1860, Namacus Amyot and Serville 1843, Nisoscolopocerus Barber 1928, Scolopocerus Uhler 1875, Sethenira Spinola 1837, Villasticoris Brailovsky 1990, Zicca Amyot and Serville 1843
Description and images of Madura perfida Stål 1862 (not yet in the guide) see here, here, here
Images of Sethenira ferruginea Stål 1870 (not yet in the guide) see here
Tribe Discogastrini Stål 1867
Tribe Leptoscelini Stål 1867
Genera Amblyomia Stål 1870, Dallacoris Osuna 1981
Description and images of Amblyomia bifasciata Stål 1870 (not yet in the guide) see here
Tribe Nematopodini Amyot and Serville 1843
Genera Mamurius Stål 1862, Mozena Amyot and Serville 1843, Piezogaster Amyot and Serville 1843, Thasus Stål 1865
Tribe Spartocerini Amyot and Serville 1843
Genera Sephina Amyot and Serville 1843, Spartocera Laporte 1833
Subfamily Meropachinae Stål 1867
Tribe Merocorini Stål 1870
Genera Merocoris Perty 1833
Subfamily Pseudophloeinae Stål 1867
Genera Ceraleptus Costa 1847, Coriomeris Westwood 1842
Ceraleptus  Coriomeris  
See Also Broad-headed Bugs, Alydidae
Print References The Century Dictionary--entries for Coreus, Coreidae (1)
Eisner, pp. 92, 234-244, includes high-resolution images of glands and associated structures (4)
Castner, p. 88, figs. 309-311 (5)
Bland and Jacques, pp. 152-153, fig. 128A (scent glands), fig. 140 (6)
Borror and White (1st ed.) pp. 122-123--good comparison with Alydidae (8)
Peterson's First Guide to Insects (9)
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