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Subfamily Campsomerinae

I love that pollen! - Dielis plumipes - male Scoliid wasp? - Dielis plumipes - female Scoliidae - Dielis plumipes - male WaspinSand - Dielis plumipes Wasp - Dielis plumipes Wasp? - Dielis pilipes - male Maryland Dielis - Dielis plumipes - male Xanthocampsomeris completa? - Xanthocampsomeris completa
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hymenoptera (Ants, Bees, Wasps and Sawflies)
No Taxon (Aculeata - Ants, Bees and Stinging Wasps)
Superfamily Scolioidea
Family Scoliidae (Scoliid Wasps)
Subfamily Campsomerinae
Synonyms and other taxonomic changes
The scoliid subfamilies used on BugGuide are alternatively represented as tribes (Campsomerini and Scoliini) united under a differently-defined Scoliinae.
Our representation of Campsomerinae has variably been sub-divided (either as tribes or subtribes). As used by Argaman (1996), these tribes would include Campsomerini (for the present genera of Dielis, Pygodasis, and Xanthocampsomeris) and Colpini (for the present genera of Colpa and Crioscolia). These subtaxa were not used by Osten (2005).(1)
Subsequent phylogenetic studies by Khouri et al. (2022) suggest establishing separate tribes of Campsomerini and Colpini, with Colpini as most sister to Scoliini. They additionally found that Dielis pilipes grouped within Xanthocampsomeris rather than dielis but did not formally transfer the species.(2)
Explanation of Names
Campsomerini Betrem, 1972
Campsomerinae Betrem, 1972
Numbers
16 spp. in 4 genera in our area; >30 genera total
Identification
Overview of Campsomerini
Separated from Scoliinae (Scoliini) by 2 veins at posterior margin of submarginal cell(3) forming a narrow, L-shaped enclosure


Overview of subtaxa
Campsomerine genera may be further divided into two groups (formerly treated as tribes or subtribes). Members of the Campsomeris group / Campsomerini s.s. (Dielis, Pygodasis, and Xanthocampsomeris) have two submarginal cells. Members of the Colpa group / Colpini (Colpa and Crioscolia) have three submarginal cells.(3)


Members of the Colpa group are separated into two subgenera, Colpa (Colpa) and Colpa (Crioscolia) based on the interantennal space in females and the antennal structure in males.
Colpa (Colpa): female interantennal space less strongly elevated, male with flagellum not modified
Colpa (Crioscolia): female interantennal space forming a strongly elevated platform, male with proximal part of flagellum conspicuously narrower than distal part

Females of the Campsomeris group in our area may be quickly separated based on the color of hair on the head (note: not the pronotal collar, which could cause confusion with Dielis plumipes). This trait is not typically useful with males. — J. Hoskins
Dielis: white hairs
Pygodasis: black hairs
Xanthocampsomeris: reddish-brown hairs
Range
Cosmopolitan below about 51°N latitude.
Remarks
Campsomeris (sensu stricto) is a Neotropical genus and does not occur in our area.
Print References
Argaman Q. (1996) Generic synopsis of Scoliidae (Hymenoptera, Scolioidea). Ann. hist.-nat. Mus. natn. hung. 88: 171–222.
Bradley, J. Chester. 1957. The taxa of Campsomeris (Hymenoptera: Scoliidae) occurring in the New World Transactions of the American Entomological Society, 83: 65–77.(4)
Bradley, J. Chester. 1964. Further notes on the American taxa of Campsomeris (Hymenoptera: Scoliidae). Entomological News, 75: 101–108.(5)
Works Cited
1.Checkliste der Dolchwespen der Welt (Insecta: Hymenoptera, Scoliidae)
von Till Osten. 2005. Bericht der Naturforschenden Gesellschaft Augsburg, 62: 1–62.
2.The evolutionary history of mammoth wasps (Hymenoptera: Scoliidae)
Ziad Khouri, Jessica .P. Gillung, & Lynn S. Kimsey. 2022. bioRxiv preprint.
3.Grissell (2013): Scoliid Wasps of Florida
4.The taxa of Campsomeris (Hymenoptera: Scoliidae) occurring in the New World
J. Chester Bradley. 1957. Transactions of the American Entomological Society, 83: 65–77.
5.Further notes on the American taxa of Campsomeris (Hymenoptera: Scoliidae)
J. Chester Bradley. 1964. Entomological News, 75: 101–108.