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Subgenus Fuscopolistes (Polistes Subgenus Fuscopolistes)
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Northern Paper Wasp (Polistes fuscatus)
Photo#1081953
Copyright © 2015
Vincent Vizachero
Wasp? -
Polistes fuscatus
Baltimore City County, Maryland, USA
June 10, 2015
Photographed near sunset on New Jersey tea (ceanothus americanus).
Images of this individual:
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Contributed by
Vincent Vizachero
on 10 June, 2015 - 6:59pm
Last updated 13 October, 2023 - 4:19pm
Fuscatus vs. metricus
I have been asked to comment. The bulge of sternum 2 is very low. That is consistent with fuscatus. The pattern of tergum 2 more resembles metricus. I am leaning more towards fuscatus overall, but even with a specimen at hand I find these two species sometimes hard to separate.
…
Matthias Buck
, 29 October, 2023 - 10:05pm
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The third iNaturalist Polistine top identifier
Chris Kratzer also thought this photo is metricus like I did, although wasn't certain. In any case, Jonathan's claim of how you would ID this record is somewhat different from what you now have said, so it was correct to ask you or additional identifiers to review. Is the head color character truly diagnostic of fuscatus on it's own? Also, for the similar fuscatus photos Jonathan shared, first I might wonder if any of those could be metricus, maybe not, but either way those photos look different than the current one.
As I also originally said, the shape of the red T1 and T2 markings are most typical of metricus, vs. he said that isn't true. Is it true that many fuscatus have red markings shaped like this? Additional comments or asking additional identifiers would be helpful. It would be interesting to know what iNaturalist pedro3111 or others would ID this as. In any case, would it be best to move this record from fuscatus to "fuscatus or metricus"?
…
bdagley
, 30 October, 2023 - 9:56am
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Patterns
For the Vespid Atlas, it was my goal to make identification of vespids as easy as possible, i.e. wherever possible I tried to identify unique colour patterns that would characterize a species. This proved to be a very difficult task in Fuscopolistes because fuscatus is so variable that it shares patterns with most other species it occurs together with. The best I could do was keying out fuscatus females in several places taking into consideration various colour morphs. I was unable to come up with a conclusive diagnosis of fuscatus vs. metricus based on colour patterns. This was backed up by DNA sequencing which I did extensively (see my Polistes paper in Zootaxa). In some cases, the boundaries of variation could only be determined by sequencing (e.g., between fuscatus and the northern form of bellicosus, which previously was confused with the former).
The pattern of T 1 and 2 shown here also occurs in fuscatus. I have a few hundred specimens from MD and surrounding states here at the museum. Every wasp has a different pattern and some are very similar to this one. I am not sure in which way the head pattern of this wasp serves to rule out metricus. What I stated back then is that the clypeus of metricus is never marked with black, but sometimes it is in fuscatus with metricus-like pattern. I still consider this to be true today.
In conclusion, I find the evidence based on coloration ambiguous. The structural character (evenly rounded S2) weighs much more significantly in my opinion. Without hearing any other opinions, I would have identified this wasp as fuscatus. But I have no problem with backing it up to 'fuscatus or metricus'. Since different people independently have come to different conclusions this is probably the best course of action.
…
Matthias Buck
, 30 October, 2023 - 11:02am
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Okay, that information is useful.
The head/face character I referred to was as commented to me on this page, "P. metricus *never* has the black markings of the face extending broadly to the antennal bases." I'm wondering if this is always the case.
On the topic of coloration, although not pertaining to this record, I've also wondered if metricus can have red markings past T2 vs. the key says never past T2.
…
bdagley
, 30 October, 2023 - 12:38pm
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Abdominal markings
Metricus never has red markings beyond T2, except in the Florida Keys. Most females there have red on T3 and occasionally even T4. To my knowledge, fuscatus does not occur in the Keys but it is found in the Everglades.
…
Matthias Buck
, 30 October, 2023 - 2:12pm
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Moved
Moved from
Metric Paper Wasp
.
Again,
P. metricus
*never*
has the black markings of the face extending broadly to the antennal bases, per Matthias Buck. As that is clearly the case here, we are not dealing with
P. metricus
. S2 is also much more convex in
P. metricus
; again, the flat area isn't long enough to use as a diagnostic toward
P. metricus
. This cleanly keys out to
p. fuscatus
in Matthias's keys.
Matthias has also noted, numerous times, populations of
P. fuscatus
in Maryland that have almost identical abdominal patterns to
P. metricus
.
Whatever "additional characters" you're finding are superficial traits that cannot be used reliably. Relying upon the wrong traits will simply confuse the species in the same way as did early authors.
…
Jonathan Hoskins
, 13 October, 2023 - 4:20pm
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I still disagree, and additional identifiers disagreed
It would be best to ask him what species he thinks this is and to share the answer here, given that we disagree.
…
bdagley
, 13 October, 2023 - 4:42pm
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Moved
Moved from
Northern Paper Wasp
.
…
bdagley
, 13 October, 2023 - 3:06pm
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Moved
Moved from
Metric Paper Wasp
.
Black markings extend below the antennal bases, so this isn't
P. metricus
. S2 also doesn't have the requisite convexity - the flat portion is noticeably shorter.
…
Jonathan Hoskins
, 12 October, 2023 - 10:35pm
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This is metricus, which additional identifiers I asked agree w/
The abdomen convexity is sufficient to indicate metricus. Another indication is the shape of the red markings on the first two segments, and there are additional characters indicating metricus in these photos.
…
bdagley
, 13 October, 2023 - 3:05pm
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Moved
Moved from
Metric Paper Wasp
.
…
bdagley
, 8 October, 2023 - 12:59pm
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Moved
Moved from
Subgenus Fuscopolistes
.
…
bdagley
, 8 October, 2023 - 12:57pm
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Moved
Moved from
Umbrella Paper Wasps
.
…
bdagley
, 10 May, 2023 - 12:02pm
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Moved
Moved from
ID Request
.
…
John S. Ascher
, 10 June, 2015 - 10:44pm
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I added a dorsal view
I'm not sure this view helps narrow it down much, but it's the best I captured of this specimen.
…
Vincent Vizachero
, 11 June, 2015 - 11:55pm
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yes
a vespid. looks like a very red
Polistes
. One of the experts will get you further
…
Steve Scholnick
, 10 June, 2015 - 9:52pm
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Thank you for helping!
I'm guessing Polistes metricus or Polistes fuscatus?
…
Vincent Vizachero
, 11 June, 2015 - 5:26am
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tough call
You're welcome. It might be hard to ID to species without a dorsal view. The fact that Dr. Ascher left it at the genus level is consistent with that. You might eventually get a species ID but that's most likely beyond me.
…
Steve Scholnick
, 11 June, 2015 - 10:00pm
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Added a dorsal view
The view is not ideal, but is the best one I captured.
…
Vincent Vizachero
, 11 June, 2015 - 11:54pm
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