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For Insects, Spiders & Their Kin
For the United States & Canada
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Family Cleridae - Checkered Beetles

Checkered Beetle - Cymatodera undulata Enoclerus muttkowskii in New Brunswick - Enoclerus muttkowskii Clerid - Cymatodera Ornate Checkered Beetle - Trichodes ornatus Clerid - Cymatodera balteata  Cleridae - Cymatodera aegra-complex Cleridae, tabida or curtipennis, dorsal - Isohydnocera Cleridae, lateral  - Enoclerus rosmarus
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga
No Taxon (Series Cucujiformia)
Superfamily Cleroidea
Family Cleridae (Checkered Beetles)
Other Common Names
Checkered Flower Beetles(1)
Synonyms and other taxonomic changes
family classification has been recently revised, with multiple new subfamilies erected(2) (the arrangement still followed here), although only 5 subfamilies are reconized in (3)
Explanation of Names
Cleridae Latreille 1802
almost 300 spp. in 36 genera in our area(4), 50 in Canada(5); >3,600 spp. in >330 genera of 12 subfamilies worldwide(2)
Nearctic fauna catalogued in(6)
Overview of our fauna classification adapted from(2); * –taxa not yet in the guide]
Subfamily ISOCLERINAE Ababa

====a much more compact classification used in(7)=========
Subfamily TILLINAE Fischer von Waldheim 1813
Subfamily HYDNOCERINAE Spinola 1844
__Tribe Hydnocerini Spinola 1844
Subfamily CLERINAE Latreille 1802
Subfamily KORYNETINAE Laporte 1836
2-24 mm(4)
Body usually long and narrow, often colorful, covered with bristly hairs; head usually wider than pronotum; pronotum sybcylindrical and narrower than elytra; antennae vary considerably and are of great diagnostic value within the group
online sources for visual identification:(8)(9)(10)(11)
worldwide, much more diverse in warmer climes; in our area more diverse in the south(4)
insects, esp. wood- and cone-borers; some adults feed on pollen; a few species are scavengers(4)
Enoclerus, Phyllobaenus, Isohydnocera, Cymatodera, and Placopterus spp. have been recorded as gall infiltrators. --J.M. Leavengood
Print References
Internet References
Works Cited
1.A Manual of Common Beetles of Eastern North America
Dillon, Elizabeth S., and Dillon, Lawrence. 1961. Row, Peterson, and Company.
2.Classification, natural history, phylogeny, and subfamily composition of the Cleridae and generic content of the subfamilies
Opitz W. 2010. Entomologica Basiliensia et Collections Frey 32: 31–128.
3.Comprehensive phylogeny of the Cleroidea (Coleoptera: Cucujiformia)
Gimmel M.L., Bocakova M., Gunter N.L., Leschen R.A.B. 2019. Sys. Entomol. 44: 527-558.
4.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). 2002. CRC Press LLC, Boca Raton, FL.
5.The checkered beetles (Coleoptera: Cleridae) of the Maritime Provinces of Canada
Majka C.G. 2006. Zootaxa 1385: 31–46.
6.Catalogue of North American beetles of the family Cleridae
Wolcott A.B. 1947. Fieldiana: Zoology 32: 63–105.
7.Family-group names in Coleoptera (Insecta)
Bouchard P, Bousquet Y, Davies A, Alonso-Zarazaga M, Lawrence JF, Lyal CH, Newton A, Reid CA, Schmitt M, Ślipiński SA, Smith A. 2011. ZooKeys 88: 1–972.
8.A guide to the Cleridae of Atlantic Canada, by Christopher Majka
9.Ayres B.D., Barr W.F. (YYYY) Photographs of North American Cleridae associated with conifers
10.Jacques Rifkind's Enoclerus Photographs
11.The checkered beetles (Coleoptera: Cleridae) of Florida.
Leavengood, Jr., J.M. 2008. University of Florida, Gainesville. 206 pp.
12.Preliminary survey of the checkered beetles (Coleoptera: Cleridae) of Tamaulipas, Mexico.
Hernández-May, M.A. et al. 2021. The Coleopterists Bulletin 75(2): 661-669.
13.The Clerid Homepage, by R. Gerstmeier