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Subfamily Rutelinae - Shining Leaf Chafers

Beyer's Scarab - Chrysina beyeri oriental for june - Anomala orientalis Shining Leaf Chafer - Anomala marginata Habitus Dorsal - Cotalpa subcribrata Popillia japonica Goldsmith Beetle? - Cotalpa lanigera Grapevine beetle - Pelidnota punctata Beetle  - Pelidnota punctata
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
Superfamily Scarabaeoidea (Scarab, Stag and Bess Beetles)
Family Scarabaeidae (Scarab Beetles)
Subfamily Rutelinae (Shining Leaf Chafers)
Explanation of Names
Author is MacLeay 1819, who originally described as its own family, Rutelidae, now regarded as a subfamily of the Scarabaeidae. Name from genus Rutela (Latreille, 1817), an error for Latin rutila (masculine rutilus), meaning red (1).
Worldwide, over 4,100 species.
Elongated to oval, often shiny, scarabs. Distinctive for having unequal, independently moveable tarsal claws, esp. on hind legs (2). Other typical (but not necessarily diagnostic) characters include:
middle tibiae with two apical spurs
pygidium exposed, not completely covered by elytra
scutellum is exposed, not covered by elytra
antennae with 9-10 segments, last three forming a club

Photos illustrating these characteristics:
Adults feed on foliage and fruit. Larvae feed on roots and decaying vegetation (2).
See Also
Cetoniinae - Fruit and Flower Chafers
Melolonthinae - May Beetles and June Bugs
Print References
The Century Dictionary--entry for Rutela (1)
White, pp. 144-147 (2)
Harpootlian, p. 97 (3)
Dillon, p. 506 (4)
Ratcliffe & Paulsen (5)
Works Cited
1.The Century Dictionary: an encyclopedic lexicon of the English language
2.Peterson Field Guides: Beetles
Richard E. White. 1983. Houghton Mifflin Company.
3.Scarab beetles (Coleoptera: Scarabaeidae) of South Carolina
Phillip J. Harpootlian. 2001. Clemson University Public Service.
4.A Manual of Common Beetles of Eastern North America
Dillon, Elizabeth S., and Dillon, Lawrence. 1961. Row, Peterson, and Company.
5.The Scarabaeoid Beetles of Nebraska
Brett C. Ratcliffe & M.J. Paulsen. 2008. University of Nebraska State Museum, Vol 22, 570 pp.