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Subfamily Sciapodinae

Unidentified Insect 45 - Condylostylus - female Tiny green fly - Condylostylus nigrofemoratus - male Longlegged Fly IMG_0208 - Condylostylus Longlegged Fly - Condylostylus - female my first bug, is it a fly or some kind of bee? - Condylostylus - male Fly ID request - Condylostylus mundus - male Condylostylus - caudatus ?? - Condylostylus caudatus - male - female Small Fly - Condylostylus occidentalis - female
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Diptera (Flies)
No Taxon (Orthorrhapha)
Superfamily Empidoidea
Family Dolichopodidae (Longlegged Flies)
Subfamily Sciapodinae
Current classification has four genera: Amblypsilopus (16 species), Condylostylus (35 species), Mesorhaga (12 species), and Sciapus (8 species).
Most Sciapodinae have vertex deeply excavated when viewed from ahead

and vein M1+2 with a long, promient fork.

Other Dolichopodidae have at most a slightly excavated vertex or a very short fork. Mesorhaga is the only genus in North America to uniformly lack the second branch of M; it may also be absent in a few males of Amblypsilopus.
Unlike most Dolichopodidae, Sciapodinae usually perch with wings spread.
Worldwide, more diverse in Southern hemisphere and tropics. Only five species west of Rocky Mountains, one introduced.(1)
Most green long-legged flies perched on broad leaves are in this subfamily.
Revised at the genus level by Bickel (1994(2)).
Keys to Sciapus s.l. (including Amblypsilopus) in Steyskal 1966, 1973.

Key to North American genera and selected species of Sciapodinae (subgeneric keys on genus pages) adapted from Aldrich 1915, Bickel (1994(2), 2009(3)), Loew 1864, Steyskal 1966 and 1973.

1. Hind femur with anterior preapical seta (sometimes absent in male); acrostichals short or absent; dorsocentrals usually long in both sexes; lateral scutellar setae short and hairlike; thorax often pruinose. Wing unmarked. Eastern North America, Washington → Sciapus
1'. Femora without strong anterior preapical setae; acrostichals usually present; thorax usually shining metallic → 2

2. Vein M2 completely absent, dorsocentrals strong on both sexes; arista usually dorsal; strong vertical seta present in both sexes, not arising from setose mound; tibiae without major setae → Mesorhaga
2'. Vein M2 present, sometimes only as a fold; other features various → 3

3. Frons of both sexes with raised setose mound bearing strong vertical seta; M1 beyond M2 usually sharply recurved basad; both pairs of scutellar setae long; wing often with dark bands, usually joined in U shape; costa of male usually without cilia; both sexes with 4-5 strong dorsocentrals; antenna black → Condylostylus
3'. Frons with vertical seta strong in female, usually weak in male, but without vertical seta arising on setose mound; males usually with some anterior dorsocentrals weak and hairlike; M1 usually with gentle arc to apex or rounded right angle bend and R5 approaching M; antenna often with yellow; costa of male often with cilia → Amblypsilopus
Print References
Steyskal, G. C. 1966. A new species of Sciapus from California, with a revised key to the species of Sciapus found in America north of Mexico (Diptera, Dolichopodidae). Proc. Ent. Soc. Washington. 68:290-294.
Steyskal, G. C. 1973. Notes on Southeastern North American Species of Condylostylus and Sciapus (Diptera:Dolichopodidae). The Florida Entomologist 56:359-360. (JSTOR)
Works Cited
1.Catalog of the Dolichopodidae (Diptera) of America north of Mexico
Marc A. A. Pollet et al. 2003. American Museum of Natural History.
2.The Australian Sciapodinae
Daniel J. Bickel. 1994. Australian Museum.
3.Manual of Central American Diptera
Brian V. Brown et al. 2009. NRC Research Press.