Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada

Family Passalidae - Bess Beetles


Schoolmeesters P. (2012) World Scarabaeidae database
[cite:890259]

Covers all Scarabaeoidea; currently accessible through Annual Checklist interface only

Beckemeyer R.J. (2003) Dung beetles & their relatives in Kansas: an annotated checklist based on a review of literature
[cite:745901]

Scarabaeoidea of southern South America
[cite:601321]

Lawrence J.F., et al. (1995-2005) Elateriformia (Coleoptera) larvae: descriptions, illustrations, identification...
[cite:747404]
Full citation: Lawrence J.F., Hastings A.M., Dallwitz M.J., Paine T.A., Zurcher E.J. 1995 onwards. Elateriformia (Coleoptera) larvae: descriptions, illustrations, identification, and information retrieval for families and subfamilies. Version: 9th October 2005.

Lawrence J.F., et al. (2000-2005) Elateriformia (Coleoptera): descriptions, illustrations, identification...
[cite:506954]
Full citation: Lawrence J.F., Hastings A.M., Dallwitz M.J., Paine T.A., Zurcher E.J. 2000 onwards. Elateriformia (Coleoptera): descriptions, illustrations, identification, and information retrieval for families and subfamilies. Version: 9th October 2005

The Beetle Ring
This site links to numerous other sites about beetles, with short descriptors of each.

Ciegler J.C. (2014) South Carolina beetles
[cite:928719]

Checklist of Coleoptera Known from Great Smoky Mountains National Park
[cite:899594

GSMNP beetle species count (as of 11 February 2013) is 2,522.

Information contained on the Excel spreadsheet includes, family, subfamily (in part), genus and species, a comment line, source(s) of the record, and references to publications dealing with GSMNP beetles.

Prepared by:
The Coleoptera Taxonomic Working Group (TWIG) at the Louisiana State Arthropod Museum (LSAM)
Christopher Carlton, TWIG Coordinator