Genus Hesperia - Branded Skippers
Phylogenetic relationships of subfamilies and circumscription of tribes in the family Hesperiidae (Lepidoptera: Hesperioidea).By Warren, A.D., J.R. Ogawa and A.V.Z. Brower.
Cladistics 24, 1–35., 2008
Warren, A.D., J.R. Ogawa and A.V.Z. Brower. 2008. Phylogenetic relationships of subfamilies and circumscription of tribes in the family Hesperiidae (Lepidoptera: Hesperioidea). Cladistics 24, 1–35.
A comprehensive tribal-level classification for the worlds subfamilies of Hesperiidae, the skipper butterflies, is proposed for the first time. Phylogenetic relationships between tribes and subfamilies are inferred using DNA sequence data from three gene regions (cytochrome oxidase subunit I-subunit II, elongation factor-1a and wingless). Monophyly of the family is strongly supported, as are some of the traditionally recognized subfamilies, with the following relationships: (Coeliadinae + ("Pyrginae" + (Heteropterinae + (Trapezitinae + Hesperiinae)))). The subfamily Pyrginae of contemporary authors was recovered as a paraphyletic grade of taxa. The formerly recognized subfamily Pyrrhopyginae, although monophyletic, is downgraded to a tribe of the "Pyrginae". The former subfamily Megathyminae is an infra-tribal group of the Hesperiinae. The Australian endemic Euschemon rafflesia
is a hesperiid, possibly related to "Pyrginae" (Eudamini). Most of the traditionally recognized groups and subgroups of genera currently employed to partition the subfamilies of the Hesperiidae are not monophyletic. We recognize eight pyrgine and six hesperiine tribes, including the new tribe Moncini.
Contributed by Mike Quinn
on 3 January, 2018 - 2:21pm
Revised classification of the family Hesperiidae (Lepidoptera: Hesperioidea) based on combined molecular and morphological data.By Warren, A.D., J.R. Ogawa & A.V.Z. Brower.
Systematic Entomology 34(3): 467-523., 2009
For a copy
Warren, A.D., J.R. Ogawa & A.V.Z. Brower. 2009. Revised classification of the family Hesperiidae (Lepidoptera: Hesperioidea) based on combined molecular and morphological data. Systematic Entomology 34(3): 467-523.
We propose a revised higher classification for the genera of Hesperiidae (skipper butterflies) of the world. We have augmented our published DNA data matrix with 49 morphological characters in order to infer relationships for taxa not sampled in the molecular study. We use the results of a combined analysis to identify morphological synapomorphies of the suprageneric clades of Hesperiidae, and to hypothesize a phylogenetic classification of the world’s genera of Hesperiidae, the first of its kind for this diverse group.
Contributed by Mike Quinn
on 3 January, 2018 - 2:10pm
Genomic evidence suggests further changes of butterfly namesBy Zhang, J., Q. Cong, J. Shen, P.A. Opler, N.V. Grishin
Taxonomic Report of the International Lepidoptera Survey, 8(7): 1-41., 2020
Zhang, J., Q. Cong, J. Shen, P.A. Opler, N.V. Grishin, 2020. Genomic evidence suggests further changes of butterfly names. Taxonomic Report of the International Lepidoptera Survey, 8(7): 1-41
Contributed by Steve Nanz
on 9 June, 2022 - 1:58pm
Butterflies of Pennsylvania, a field guideBy James L. Monroe, David M. Wright
University of Pittsburgh Press
From the publishers page:
This work has all of the features that make field guides to a region's butterfly fauna useful to anyone with a serious interest in that fauna. . . . the book is a bargain and a must for anyone with an interest not just in Pennsylvania's fauna, but the northeast fauna as a whole.
News of the Lepidopterists' Society
Winner, 2017 National Outdoor Book Award
Though I am in the southeast, I own this field guide and find it very useful - Roy Cohutta Brown.