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Smilax leaf blister galls - Meunieriella on-smilax

Smilax leaf blister galls - Meunieriella on-smilax
Upper Iowa River WMA, Bronner Tract, Winneshiek County, Iowa, USA
September 24, 2015
Greenbriar, Smilax tamnoides, NOT Smilax rotundifolia, as per below

Host plant is Smilax tamnoides
The host plant in this image is actually Smilax tamnoides. The identifying needle-shaped bristly prickles are visible on the stem.

Thank you.

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