Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada

Subfamily Scolytinae - Bark and Ambrosia Beetles

Bark beetle - Phloeotribus liminaris small beetle with mites - Xyleborus celsus Hylocurus sp. - Hylocurus spadix Pseudohylesinus nebulosus Beetle - Dryocoetes betulae - female Xyloterinus politus (?) - Xyloterinus politus Xylosandrus crassiusculus dorsal view - Xylosandrus crassiusculus Pseudopityophthorus pruinosus - male
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
No Taxon (Series Cucujiformia)
Superfamily Curculionoidea (Snout and Bark Beetles)
Family Curculionidae (Snout and Bark Beetles)
Subfamily Scolytinae (Bark and Ambrosia Beetles)
Other Common Names
Engraver Beetles
Synonyms and other taxonomic changes
used to be treated as a separate family, Scolytidae (=Ipidae); all the subtribes used in the guide (except Corthylina & Pityophthorina) are treated as tribes in(1)(2)
Explanation of Names
Scolytinae Latreille 1804
~560 spp. in >100 genera in our area(3), ~6000 spp. in almost 250 genera worldwide arranged into ~30 [sub]tribes(1)(4)(2)
most comprehensive treatment of NA fauna in(5)
illustrated online keys to US genera(6), se. US spp.(3), LA spp.(7), TX+OK spp.(8), KS spp.(9)
invasive spp. discussed in(10)(11)
fungi, mites, and nematodes vectored by nonindigenous bark- or wood-boring beetles established in Canada listed in(12)
Works Cited
1.Handbook of Zoology. Coleoptera Vol. 3: Morphology and Systematics (Phytophaga)
Leschen R.A.B., Beutel R.G. (eds). 2014. Walter de Gruyter, Berlin. 687 pp.
2.Family-group names in Coleoptera (Insecta)
Bouchard P. et al. 2011. ZooKeys 88: 1–972.
3.Atkinson T.H. (200_-2018) Bark and ambrosia beetles
4.Bark beetles: biology and ecology of native and invasive species
Vega F.E., Hofstetter R.W. (eds.). 2015. Elsevier. 620 pp.
5.The Bark and Ambrosia Beetles of North and Central America (Coleoptera: Scolytidae), a Taxonomic Monograph
Stephen Wood. 1982. Great Basin Naturalist Memoirs, Number 6, Brigham Young University.
6.Mercado J.E. (2011) Bark beetle genera of the United States
7.Ferro M.L. (2006) The bark and ambrosia beetles (Curculionidae: Scolytinae) of Louisiana: an interactive web-based key
8.Atlas and checklist of the bark and ambrosia beetles of Texas and Oklahoma (Curculionidae: Scolytinae and Platypodinae)
Atkinson T.H., Riley E.G. 2013. Insecta Mundi 02920: 1-46.
9.Guide to the bark and ambrosia beetles of Kansas
Salsbury G.A. 2004. Kansas Dept. Agric., Topeka KS. 26 pp.
10.Exotic bark and ambrosia beetles (Coleoptera: Curculionidae: Scolytinae) in the United States: Current and potential invaders
11.Exotic bark and ambrosia beetles (Coleoptera: Curculionidae: Scolytinae) in the United States: potential and current invaders
Haack R.A., Rabaglia R.J. 2013. In: Potential invasive pests of agricultural crops. Peña J.E., ed. CABI International, Wallingford, UK, pp. 48-74.
12.Forest biosecurity: alien invasive species and vectored organisms
Humble L.M., Allen E.A. 2006. Can. J. Plant Pathol. 28: S256–S269.