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Photo#1466055
Extra Cross-vein Location - Pygodasis quadrimaculata - male

Extra Cross-vein Location - Pygodasis quadrimaculata - Male
Florida, USA
The males of this species may have this cross-vein and the females do not have it. This vein subdivides the second discal cell, directly underneath the second sub-marginal cell.
Image adapted from this post by Jeff HollenBeck: Compare with the wings of a female: