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Family Behningiidae - Sand-burrowing Mayfly

Dolania americana
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Ephemeroptera (Mayflies)
Suborder Furcatergalia
Family Behningiidae (Sand-burrowing Mayfly)
6 spp. in 3 genera total(1)(2)
Medium-sized to large mayfly
Nymph: somewhat flattened but otherwise highly unusual, adapted for burrowing in sand; mandibles without tusks; head and prothorax with dorsal pad of long spines on each side; forelegs palplike, middle- and hind legs modified as protective devices for gills; gills ventral
key to spp. in(2)
Holarctic, with 2 genera in Eurasia and one in North America(1)
Mid-sized to large rivers with sand bottoms; nymphs usually in clean sand
Nymphs are predators; adults are short-lived and non-feeding
Life Cycle
North American populations univoltine