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Species Aphodius granarius - Aphodius (Calamosternus) granarius

Scarabaeidae - ? - Aphodius granarius Aphodiine scarab 4 - Aphodius granarius Aphodiine scarab 4 - Aphodius granarius Dung Beetle - Aphodius granarius Aphodiine Dung Beetle? - Aphodius granarius Scarab: Aphodius granarius? - Aphodius granarius Aphodiine - Aphodius granarius Small scarab under rock - Aphodius granarius
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
Superfamily Scarabaeoidea (Scarab, Stag and Bess Beetles)
Family Scarabaeidae (Scarab Beetles)
Subfamily Aphodiinae (Aphodiine Dung Beetles)
Tribe Aphodiini
Genus Aphodius
No Taxon (Subgenus Calamosternus)
Species granarius (Aphodius (Calamosternus) granarius)
Explanation of Names
Orig. Comb: Scarabaeus granarius Linnaeus, 1767
3-5 mm (1)
4–6 mm - Smith & Skelley, 2007

A highly variable species as indicated by the large synonymy.
Males have broad pronotum and smoother clypeus;
females have narrower pronotum and rugose clypeus. - Smith & Skelley, 2007
Cosmopolitan, one of the most widely dist. sp., now found in every part of the globe. (1)
One of the most common sp. in cow dung in the ne. US. (1)
common wherever cows and other European farm animals are found - Smith & Skelley, 2007
Highly polyphagous, feeding on all kinds of dung, debris, decaying vegetables, compost, carrion, etc. (1)
Type Locality: "Europa"
Internet References
University of Nebraska - Andrew Smith and Paul Skelley, 2007
Works Cited
1.The Scarab Beetles of Florida
Robert Woodruff. 1973. Florida Department of Agriculture and Consumer Services.