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Subgenus Hypherpes

Pterostichus amethystinus California Coastal Dune Beetle - Pterostichus menetriesii Carabid Beetle - Pterostichus Pterostichus castaneus Pterostichus - to sp.? - Pterostichus tristis - female Ground Beetle - Pterostichus illustris Pterostichus tristis Pterostichus protractus LeConte 1860 - Pterostichus protractus - female
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Adephaga (Ground and Water Beetles)
Family Carabidae (Ground Beetles)
Subfamily Harpalinae
Supertribe Pterostichitae
Tribe Pterostichini (Woodland Ground Beetles)
Genus Pterostichus
No Taxon Subgenus Hypherpes
Synonyms and other taxonomic changes
ca. two dozen spp. have been recently moved to the reinstated subgenera Anilloferonia and Leptoferonia(1)
Explanation of Names
Hypherpes Chaudoir 1838
NA's most speciose subgenus of ~60 described and multiple undescribed spp., all in our area and arranged into 22 groups(1)(2)
all but 2 spp. are western (west coast to the Rockies), P. adoxus & P. tristis are eastern (se. Canada to NC-TN)(3)
Works Cited
1.Phylogeny and classification of Hypherpes auctorum (Coleoptera: Carabidae: Pterostichini: Pterostichus)
K.W. Will & A.S. Gill. 2008. Annals of Carnegie Museum. 77(1): 93-127.
2.Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico
Bousquet Y. 2012. ZooKeys 245: 1–1722.
3.American Beetles, Volume I: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia
Arnett, R.H., Jr., and M. C. Thomas. (eds.). 2000. CRC Press LLC, Boca Raton, FL.