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Photo#1766768
Cecidomyiidae, galls on leaves of Oak - Polystepha pilulae

Cecidomyiidae, galls on leaves of Oak - Polystepha pilulae
Whiterock Conservancy, Guthrie County, Iowa, USA
August 21, 2019
upper side of leaf

ID based on this:
and
"Circular ... evident only on upper side of leaf ... This is one of the most conspicuous Polystepha galls and is commonly seen in autumn on many kinds of oaks."
The Plant-Feeding Gall Midges of North America, Raymond Gagné

Given to me by a participant in the Asteraceae Workshop.

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Cecidomyiidae, galls on leaves of Oak - Polystepha pilulae Cecidomyiidae, galls on leaves of Oak - Polystepha pilulae