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Scarab 3 - Acoma mixta - male

Scarab 3 - Acoma mixta - Male
Picture Rocks, NW of Tucson, Pima County, Arizona, USA
May 31, 2008
Size: 4 mm
I think this is the same species that Phillip tentatively IDed as Acoma. I believe it's a male, because I found a similar one with smaller antennae

Images of this individual: tag all
Scarab 3 - Acoma mixta - male Scarab 3 - Acoma mixta - male

this looks to be Acoma mixta Howden based on the clypeus, darker color (than arizonica and rufula), and range.

Moved from Acoma.

an Acoma sp., will have to dig out the literature, probably not many species to choose from in AZ [only 4?], appear to have limited allopatric [nonoverlapping geographic areas] distributions - common in species where one or both sexes are flightless. None are evidently commonly collected. According to American Beetles, the females are still unknown, but are assumed flightless - maybe you can find one? Perhaps they're like Pleocoma - the females are significantly larger and usually need to be dug up. According to Cazier [quick glance] - since its dark brown, I'd say its probably Acoma brunnea. Acoma are currently placed in the Scarabaeidae:Melolonthinae:INCERTAE SEDIS

Acoma Casey, 1889

Acoma arizonica Brown, 1929 - US [AZ,CA?]
Acoma brunnea Casey, 1889 - US [AZ,TX]
Acoma conjuncta Howden, 1962 - US [?]
Acoma diminiata Howden, 1958 - US [?]
Acoma glabrata Cazier, 1953 (=nigrita) - US [CA], Mexico
Acoma knulli Howden, 1958 - US [?]
Acoma mixta Howden, 1958 - US [?]
Acoma rufula Howden, 1958 - US [AZ]

Acoma cazieri Saylor, 1948 - Mexico
Acoma confusa Van Dyke, 1928 - Mexico
Acoma dilemma Saylor, 1948 - Mexico
Acoma evansi Howden, 1962 - Mexico
Acoma gibsoni Howden, 1962 - Mexico
Acoma granulifrons Howden, 1958 - Mexico
Acoma incognita Howden, 1958 - Mexico
Acoma leechi Cazier, 1953 - Mexico
Acoma martini Howden, 1962 - Mexico
Acoma mimica Howden, 1962 - Mexico
Acoma minuta Cazier, 1953 - Mexico
Acoma ochlera Howden, 1958 - Mexico
Acoma parva Howden, 1958 - Mexico
Acoma robusta Van Dyke, 1928 - Mexico
Acoma rossi Saylor, 1948 - Mexico
Acoma seticollis Howden, 1958 - Mexico
Acoma sexfoliata Saylor, 1948 - Mexico
Acoma stathamae Cazier, 1953 - Mexico


Cazier, M.A. 1953. A review of the scarab genus Acoma (Coleoptera, Scarabaeidae). American Museum Novitates 1624:1-13. [available on-line, right click, download file, then open -]

Howden, H.F. 1958. Species of Acoma Casey having a three-segmented antennal club (Coleoptera: Scarabaeidae). Canadian Entomologist 90(7):377-401.

Howden, H.F. 1962. New Species and Synonymy in Acoma (Coloptera: Scarabaeidae). Canadian Entomologist 94(11):1147-1155.

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