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Subfamily Paussinae

Primitive Bombardier Beetle - Metrius contractus cool carabid - Goniotropis kuntzeni Metrius contractus  - Metrius contractus Bombardier Beetle - Metrius contractus Metrius contractus Eschscholtz, 1829 - Metrius contractus Pachyteles parca LeConte - Goniotropis parca Metrius contractus Ground beetle - Metrius contractus
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Adephaga (Ground and Water Beetles)
Family Carabidae (Ground Beetles)
Subfamily Paussinae
Seven species in two tribes are known from North America. (1)
Worldwide with 724 species in 34 genera. (1)

Ozaena - one species known from AZ
Black, convex beetles with a smooth, dull surface found from California to British Columbia, as well as Idaho.
Both spurs of front tibia apical (as opposed to one being preapical)
Head with one setigerous puncture over each eye.
Hind coxae not contiguous.
Middle coxal cavities disjunct.
Anterior coxal cavities closed.

Physea, Pachyteles, & Ozaena
Arizona to Texas
Prominant apical ridge on elytra that projects outward (called flange of Coanda). Seen well here.
Both spurs of front tibia apical (as opposed to one being preapical).
Antennomeres moniliform, thickened.
Anterior coxal cavities closed.
Hind coxae not contiguous.
Middle coxal cavities disjunct.
Metepimeron distinct.

Paussinae larvae
Key to genera based on larvae in (2)
In our area, from British Columbia, Idaho, & California to Arizona & Texas. (1)
Myrmecophily is common in this subfamily. In our area, Physea is the only myrmecophile.
Defensive chemicals are quinone-based and delivered at hot temperatures, discharged either by popping or oozing. (1)
Print References
Works Cited
1.American Beetles, Volume I: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia
Arnett, R.H., Jr., and M. C. Thomas. (eds.). 2000. CRC Press LLC, Boca Raton, FL.
2.Description and behaviour of Goniotropis kuntzeni larvae (Coleoptera: Carabidae: Paussinae: Ozaenini) and a key to....
Moore, W. and A. Di Giulio. 2006. Zootaxa 1111: 1–19.
3.The Middle American genera of the tribe Ozaenini with notes about the species in the southwestern United States and...
Ball, G.E. and McCleve, S. 1990. Quaestiones Entomologicae, 26: 30-116.