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For Insects, Spiders & Their Kin
For the United States & Canada

Genus Zopherus


False darkling beetles (Coleoptera: Melandryidae) and allies of the Atlantic Maritime Ecozone
By Majka C.G., Pollock D.A.
NRC Research Press. Ottawa, Ontario, 2010
In: Assessment of Species Diversity in the Atlantic Maritime Ecozone. Edited by: McAlpine D.F., and Smith I.M. NRC Research Press. Ottawa, Ontario. pp 453-463.
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The chapter treats Melandryidae, Tetratomidae, Scraptidae, and Synchroidae.

A contribution to the phylogeny of the Ciidae and its relationships with other Cucujoid and Tenebrionoid beetles
By Buder, G., C. Grossmann, A. Hundsdoerfer & K.-D. Klass
Arthropod Systematics & Phylogeny, 66(2): 165–190, 2008

Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia)
By Robertson J.A., Ślipiński A., Moulton M., Shockley F.W., Giorgi A., Lord N.P., McKenna D.D., Tomaszewska W., Forrester J. ...
Syst. Entomol. doi: 10.1111/syen.12138, 2015

Synopsis of adventive species of Coleoptera (Insecta) recorded from Canada. Part 4: Scarabaeoidea... [through Cleroidea]
By Klimaszewski J, Langor D, Smith A, Hoebeke E, Davies A, Pelletier G, Douglas HB, Webster RP, Bourdon C, Borowiec L, Scudder GGE
Pensoft Series Faunistica 116: 1-215, 2017
Full title: Synopsis of adventive species of Coleoptera (Insecta) recorded from Canada. Part 4: Scarabaeoidea, Scirtoidea, Buprestoidea, Byrrhoidea, Elateroidea, Derodontoidea, Bostrichoidea, and Cleroidea

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Handbook of Zoology. Coleoptera Vol. 3: Morphology and Systematics (Phytophaga)
By Leschen R.A.B., Beutel R.G. (eds)
Walter de Gruyter, Berlin. 687 pp., 2014

Review of the American Corylophidae, Cryptophagidae, Tritomidae and Dermestidae, with other studies
By T. L. Casey
New York Entomological Society, 1900
Journal of the New York Entomological Society, 8(2): 51-172, 1900

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The Bothrideridae and Colydiidae of America north of Mexico (Coleoptera: Clavicornia and Heteromera)
By Stephan K.H.
Florida Dept. of Agriculture and Consumer Services, Division of Plant Industry. xii, 65 p., 1989
Occasional Papers of the Florida State Collection of Arthropods, Vol.6
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Beetles of Western North America.
By Evans, A.V.
Princeton University Press, 2021
Publisher’s page

From the author of the acclaimed Beetles of Eastern North America, a triumphant follow-up guide to western beetles, lavishly illustrated with more than 1,500 stunning color photographs.

• The only comprehensive color photographic guide to the region’s beetles
• Covers 1,428 species from all 131 families west of the Continental Divide, and offers tips for identifying more than 450 additional species
• Features more than 1,500 stunning color photographs