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Species Bembidion lacunarium

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Thirty ground beetles new to Virginia, and a milestone
By R.L. Hoffman, S.M. Roble, R.L. Davidson
Banisteria 27:16-30, 2006
This publication is most notable for including R.L. Davidson's invaluable notes on identifying various Tachyines, especially those of the genus Paratachys.
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Species delimitation, classical taxonomy & genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae)
By Maddison, DR & JS Sproul
Zoological Journal of the Linnean Society, XX, 1 - 46, 2020
ABSTRACT: The western North American genus Lionepha is shown to contain at least 11 species through a combination of eight-gene species delimitation analyses and morphological study. In order to confirm the names of several species, we sequence DNA of primary types of several names, including a LeConte lectotype collected in the 1850s, using next-generation sequencing. We examine chromosomes of eight species, and show that all have 12 pairs of autosomesand an X0/XX sex-chromosome system. The following species are described as new: Lionepha australerasa, L. kavanaughi, L. lindrothi and L. tuulukwa. The name Lionepha erasa is shown to belong to a relatively rare, western species ranging from Oregon through Alaska; the common, widespread species previously known as Lionepha erasa now takes the name L. probata. Bembidion chintimini, B. lindrothellus and B. lummi are synonymized with L. erasa. We provide tools to identify specimens to species, including illustrations, diagnoses and distribution maps. DOI: 10.1093/zoolinnean/z1z167.

Unifying systematics & taxonomy: Nomenclatural changes to Nearctic tiger beetles based on phylogenetics/morphology/life history
By Duran, DP & HM Gough
Insecta Mundi 0727: 1- 12, 2019
Full title: Unifying systematics and taxonomy: Nomenclatural changes to Nearctic tiger beetles (Coleoptera: Carabidae: Cicindelinae) based on phylogenetics, morphology and life history.

Abstract: The taxonomy of Nearctic tiger beetles (Coleoptera: Carabidae: Cicindelinae) is reviewed in light of modern systematics research. Despite decades of published molecular phylogenies, the taxonomic nomenclature has not been formally updated since the 1950s. We generated a maximum likelihood phylogenetic tree based on three mitochondrial gene fragments (16S, COX3 and CytB) to address the placement of Nearctic taxa that were not included in recent studies; these species were historically contained within Cylindera Westwood, 1831, a polyphyletic genus. Here we describe Parvindela Duran and Gough, new genus, and propose 20 new combinations based on a plurality of data, including our topology, prior molecular phylogenetic studies, morphology and ecology: Apterodela unipunctata (Fabricius, 1775) new combination; Brasiella praecisa (Bates, 1890) new combination; Brasiella viridisticta (Bates, 1881) new combination; Cicindela amargosae (Dahl, 1939) new combination; Cicindela senilis (G.Horn, 1866) new combination; Cicindela willistoni (LeConte, 1879) new combination; Eunota californica (Menetries, 1883) new combination; Eunota circumpicta (LaFerte, 1841) new combination; Eunota fulgoris (Casey, 1913) new combination; Eunota gabbii (G. Horn, 1866) new combination; Eunota pamphila (LeConte, 1873) new combination; Eunota praetextata (LeConte, 1854) new combination; Eunota severa (LaFerte, 1841) new combination; Eunota striga (LeConte, 1875) new combination; Parvindela debilis (Bates, 1890) new combination; Parvindela celeripes (LeConte, 1848) new combination; Parvindela cursitans (LeConte, 1860) new combination; Parvindela terricola (Say, 1824) new combination; Parvindela nephelota (Bates, 1882) new combination; Parvindela lunalonga (Schaupp, 1884) new combination.

Range update for eight species of the subfamily Elaphrinae (Coleoptera: Carabidae) in Ontario's Far North & Nunavut, Canada
By Fleming, KJ & Beresford, DV
The Coleopterists Bulletin 73(2): 433 - 439, 2019
Title abbreviated.

ABSTRACT: Adults of eight species of beetles in the subfamily Elaphrinae (Coleoptera: Carabidae: Elaphrinae) were collected in the Far North of Ontario and on Akimiski Island, Nunavut, Canada. We found first records for Nunavut (territory) of seven species: Blethisa julii LeConte, 1863; Blethisa quadricollis Haldeman, 1847; Diacheila arctica (Gyllenhal, 1810); Elaphrus americanus Dejean, 1831; Elaphrus lecontei Crotch, 1876; Elaphrus clairvillei Kirby, 1837; and Elaphrus fuliginosus Say, 1830. Our record of one species, Elaphrus californicus Mannerheim, 1843, is a new range record for the province of Ontario.

Carabid Beetles: Their Evolution, Natural History, and Classification
By Erwin TL, Ball GE, Whitehead DR, Halpern AL (editors)
Proceedings of the First International Symposium of Carabidology, 1976

Eight Ground Beetles (Coleoptera: Carabidae) New to Virginia, with Additional Records for West Virginia and Maryland
By Curt W. Harden
Banisteria No. 50: 15-20, 2018
Abstract: Eight ground beetle species, including Trechus quadristriatus (Schrank), Elaphropus dolosus(LeConte), Brachinus americanus (LeConte), Pterostichus tenuis (Casey), Cyclotrachelus convivus (LeConte), Agonum mutatum (Gemminger & Harold), Somotrichus unifasciatus (Dejean), and Cymindis elegans LeConte, are reported from Virginia for the first time, and one species, Clinidium apertum allegheniensis Bell & Bell, is removed from the state list. Two species are reported as new state records for Maryland ([I]Harpalus katiae

Catalogue of Palaearctic Coleoptera Volume 1 (2nd edition)
By Löbl, I. & Löbl, D.
Brill , 2017
This second updated edition of the first volume (1st ed. published 2003) is a complete catalogue of suborders Archostemata, Myxophaga, and Adephaga of the Palaearctic faunal region. The latter suborder is the largest of the three which incorporates what western authors designate Geadephaga = Caraboidea = ground beetles sensu lato. This massive 1477-page work contains information on 33,914 taxa (together with synonyms), and increases the number of included species and other taxa by almost 5,000. In addition, thousands of species have their distributional data completed, and their ranks, systematic positions and nomenclature corrected. Almost two hundred new acts fix systematics and nomenclature, and numerous problems are discussed. Even such well known genera as Calosoma and Carabus, or tribes as Bembidiini and Panagaeini, are completely reorganized compared to the previously published catalogues. Thus, the work is a scaffold for biotic surveys, ecological studies, nature conservation, and it provides background checks on Palaearctic species introduced into other faunal regions. Subsequent volumes in this series cover the remaining Coleoptera groups.

Three carabid beetles (Coleoptera:Carabidae) new to Maryland & a preliminary annotated checklist for Cove Point, Calvert County
By Brent W. Steury & Peter W. Messer
The Maryland Entomologist, 2017
This article is available online.

Abstract: Sixty-nine carabid species (Coleoptera: Carabidae) in 37 genera and 19 tribes were documented from Cove Point, Calvert County, Maryland, during surveys from 2010 to 2016. Three species, Anisodactylus haplomus Chaudoir, Pterostichus permundus (Say), and Stenocrepis mexicana (Chevrolat) are documented for the first time from Maryland.

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