A few miles southeast of Kyburz, near China Flat Campground, El Dorado County, California, USA
June 25, 2018
[For full-size image, click this link...then click the image again after it loads in your browser window.]
Found on flowers of
Darmera peltata growing at the edge of a creek in a montane mixed-coniferous forest of the central Sierra Nevada.
Using the
USDA "idtools.org" interactive key for sawflies, selecting just two antennal characters...namely:
1) "antennae clavate...only slightly expanded towards the apex" (i.e. not "remarkably clubbed"); and
2) "number of antennal segments 17-22" (cf. reference image here and the 2nd image in this series)
...narrows things down to one of
Calameuta,
Cephus, or
Trachelus. Remarkably, trying most of the other characters in that interactive key would not seem to separate these three genera.
But according to the 3 preceding links, only 5 species in those genera are recorded from North America:
Calameuta clavata,
Calameuta middlekauffi,
Cephus cinctus,
Cephus pygmaeus and
Trachelus tabidus. Note that
Ca. middlekauffi was described relatively recently in Smith & Schiff
(1)(2005)...and the 4 other earlier-named species were treated as members of either Tribe Cephini in Middlekauff
(2)(1969); or simply as members of the genus
Cephus in the older Ries
(3)(1937)...see
here.
I found it
not entirely straightforward to work these photos through the keys in the above references...but by cross-referencing between the descriptions, figures, and keys in those papers I arrived at
Cephus cinctus...based mainly on the ventral view of the distal portion of the abdomen in last photo of this series. In particular, the distal edge of the 8th sternite is seen there to be more-or-less
straight...as specified in
couplet 4 of the key in Ries
(3)(1937) and shown in his
Fig. 5.
However, the distal edge of the 8th sternite is
notched in:
A) Calameuta clavata...see couplet 1) of the key on pg. 9 of Middlekauff(2)(1969); Fig. 9 in Ries(3)(1937); and the ventral shots below:
B) Calameuta middlekauffi...see the last sentence of the description of the male on pg. 865 of Smith & Schiff(1)(2005);
C) Cephus pygmaeus...per couplet 4) and Fig. 4 of in Ries(3)(1937). However, this seems contradicted by the key in Middlekauff(2)(1969)...where pygmaeus and cinctus both come out at the 2nd lead of couplet 1), which indicates no notch on the posterior edge of the 8th sternite! Nevertheless, I think pygmaeus can be eliminated here, since the keys in both Middlekauff and Ries indicate it has costa & stigma uniformly brown, whereas cinctus has costa yellow and stigma black...as seen in the photos of this series.
Based on
Fig. 10 in Ries
(3)(1937) it appears the 8th sternite is also somewhat notched in
Trachelus tabidus. But regardless of that,
T. tabidus has distinctive spine-filled cavities on sternites 7 & 8...which are
not evident in my last photo here.
So, in summary,
Cephus cinctus appears to be the ID here.