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Genus Bembidion

B. quadrimaculatum? - Bembidion quadrimaculatum Bembidion (Trichoplataphus) rolandi Fall, 1922  - Bembidion rolandi Carabidae - Bembidion lakeshore carabid - Bembidion Tecopa J17 - Bembidion Bembidion planatum Bembidion semipunctatum - male Bembidion sp.? - Bembidion
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Adephaga
Family Carabidae (Ground Beetles)
Subfamily Trechinae
Tribe Bembidiini
Subtribe Bembidiina
Genus Bembidion
Synonyms and other taxonomic changes
Amerizus is now treated as a separate genus(1)
Within Bembidion, there are three major clades: the Bembidion, Odontium, and Ocydromus series(2)
Explanation of Names
Bembidion Latreille 1802
~275 spp. in our area(3) (incl. 8 adventive) arranged in 37 subgenera and species-groups, >1400 spp.(4) in >100 subgenera & species-groups worldwide(1)
Subgenera not yet in the Guide, per(1):
Actedium (B. lachnophoroides) Whitefox River drainage (SK‒AB)
Lindrochthus (B. wickhami) OR‒CA dorsal habitus
2‒8 mm
Worldwide; ¾ of the spp. in the Northern Hemisphere(1)
Prefer open, wet habitats. Highest diversity on river banks.
Adults of most spp. are active in daylight(5)
Internet References
Genus page on ToL
Works Cited
1.Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico
Bousquet Y. 2012. ZooKeys 245: 1–1722.
2.Phylogeny of Bembidion and related ground beetles (Coleoptera: Carabidae: Trechinae: Bembidiini: Bembidiina)
Maddison D.R. 2012. Mol. Phylogenet. Evol. 63: 533‒576.
3.Re-collected after 55 years: a new species of Bembidion (Coleoptera, Carabidae) from California
Maddison D.R., Sproul J.S., Will K. 2023. ZooKeys 1156: 87–106.
5.American Beetles, Volume I: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia
Arnett, R.H., Jr., and M. C. Thomas. (eds.). 2000. CRC Press LLC, Boca Raton, FL.