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Genus Aleiodes - Common Mummy Wasps

mummy - Aleiodes Braconidae - Braconid Wasps Genus Aleiodes - Mummy-wasps - Aleiodes - female Aleiodes nolophanae dorsal - Aleiodes nolophanae - male wasp - Aleiodes Aleiodes terminalis Black-winged orange ichneumonoid - Aleiodes politiceps braconid - Aleiodes malacosomatos - male Wasp - Aleiodes politiceps
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hymenoptera (Ants, Bees, Wasps and Sawflies)
No Taxon ("Parasitica" - Parasitoid Wasps)
Superfamily Ichneumonoidea (Braconid and Ichneumonid Wasps)
Family Braconidae (Braconid Wasps)
Subfamily Rogadinae (Caterpillar Mummy Wasps)
Genus Aleiodes (Common Mummy Wasps)
Numbers
at least 155 spp. in North America including Mexico; at least 212 spp. in the New World; between 200-300 spp. worldwide — by far the most diverse Rodaginae genus in the world

Nearctic species are divided into 15 species-groups (at least 1 sp., A. buoculus, without reference to a species-group):
1. albitibia-group (1 sp.)(1)
2. apicalis-group (6 spp.)(2)
3. circumstriptus/gastritor-group (11 spp.)(3)
4. compressor-group (2 spp.)(4)
5. coxalis-group (13 spp.)(5)(6)
6. gasterator-group (10 spp.)(7)(6)
7. gressitti-group (1 sp.)(4)
8. melanopterus-group (5 spp.)(8)
9. pallidator-group (5 spp.)(9)
10. pilosus-group (67 spp.)(10)
11. praetor-group (3 spp.)(1)
12. procerus-group (2 spp.)(4)
13. pulchripes-group (8 spp.)(11)
14. rugulosus-group (1 sp.)(12)
15. seriatus-group (7 spp.)(13)
16. ufei-group (1 sp.)(4)
16. unipunctator-group (3 spp.)(7)
Size
4.0-8.0 mm
Identification
Can be determined by
cyclostome "mouth." Also see this image.. As can be seen from these 2 images, the cyclostome mouth is defined by the dorso-ventrally short clypeus and the concave labrum, bordered ventrally by the mandibles.
Mummification of host remains. Here and here are examples of rogadine mummies.
Occipital carina present. (See p. 99 in Fortier 2009 (PDF)(10) for image of occipital carina.
Koinobiont endoparasitoid life cycle (See parasitoid life cycles).
Range
Worldwide, with the greatest species diversity in northern Europe, Asia, and North America(12)
Food
Attack and kill various species of leaf-feeding caterpillars; hosts include many important forest pests, such as the spongy moth, eastern tent caterpillar, forest tent caterpillar, fall webworm, tussock moths, dagger moths, prominents, cutworms, and loopers(12)
Life Cycle
larval koinobiont endoparasitoids. As with other rogadines, Aleiodes larvae mummify the host when they finish feeding on it and kill it, and pupate inside the mummy.
Remarks
A significant number of species have been described by Dr. Scott R. Shaw and Dr. Paul M. Marsh, both of whom generously helped BugGuide with braconid wasp identification, and also by Dr. Joe Fortier, who described 71 new species in a new subfamily.(10).
Internet References
          *** Wing vein morphology and general morphology of Aleiodes are found on pp. 98-99. ***
Works Cited
1.Revision of North American Aleiodes Wesmael (Part 4): the albitibia (Herrich-Schaeffer) and praetor (Reinhard) species-groups
Shaw SR, Marsh PM, Fortier JC. 1998. Proceedings of the Entomological Society of Washington, 100(3): 553-565.
2.Revision of North American Aleiodes Wesmael (Part 2): the apicalis (Brulle) species-group in the New World
Shaw, SR, Marsh PM, Fortier JC. 1998. Journal of Hymenoptera Research, 7(1): 62-73.
3.Cladistics of the Aleiodes lineage of the subfamily Rogadinae (Hymenoptera: Braconidae)
Fortier, J.C. & Shaw, S.R. 1999. Journal of Hymenoptera Research, 8: 204-237.
4.Revision of North American Aleiodes Wesmael (Part 7): the compressor (Herrich-Schaeffer), ufei (Walley), gressitti...
Marsh PM, Shaw SR. 2003. Proceedings of the Entomological Society of Washington, 105(3): 698-707.
5.Revision of Nearctic Aleiodes Wesmael (Part 8): the coxalis (Spinola) species-group (Hymenoptera: Braconidae, Rogadinae)
Shaw SR, Marsh PM, Fortier JC. 2006. Zootaxa 1314:1-30.
6.Aleiodes (Hymenoptera, Braconidae, Rogadinae) diversity in Washington U.S.A. including three new species
Joseph Fortier. 2021. Zootaxa, 4999(2): 132–142.
7.Revision of North American Aleiodes Wesmael (Part 6): the gasterator (Jurine) and unipunctator (Thunberg) species-groups
Marsh PM, Shaw SR. 2001. Proceedings of the Entomological Society of Washington, 103(2): 291-307.
8.Revision of North American Aleiodes Wesmael (Part 5): the melanopterus (Erichson) species-group
Marsh PM, Shaw SR. 1999. Journal of Hymenoptera Research, 8(1): 98-108.
9.Revision of North American Aleiodes (Part 9): the pallidator (Thunberg) species-group with description of two new species
Shaw SR, Marsh PM, Talluto MA. 2013. Zootaxa 3608 (3): 204–214.
10.A Revision of the Tetrasphaeropyx Ashm. Lineage of the Genus Aleiodes Wesmael (Hymenoptera: Braconidae: Rogadinae)
Fortier, Joseph C. 2009. Magnolia Press.
11.Revision of North American Aleiodes Wesmael (Part 1): the pulchripes Wesmael species-group in the New World
Scott R. Shaw, Paul M. Marsh, & Joseph C. Fortier. 1997. Journal of Hymenoptera Research, 6: .
12.Aleiodes wasps of eastern forests: A guide to parasitoids and associated mummified caterpillars
Shaw, Scott R. 2006. Morgantown: Forest Health Technology Enterprise Team.
13.Revision of North American Aleiodes Wesmael (Part 3): the seriatus (Herrich-Schaeffer) species-group
Marsh PM, Shaw SR. 1998. Proceedings of the Entomological Society of Washington, 100(3): 395-408.